Shrubs or trees, 10–50(–80) dm. Young stems with tightly coiled hairs. Leaves: stipules (3–)4–6(–10) mm; blade oblong-obovate or-elliptic, obovate, ovate, or suborbiculate, (8–)12–50(–78) × (4–)8–35(–64) mm, subcoriaceous to coriaceous, margins serrate or dentate or crenate in distal 1/5–3/4, abaxial veins with coiled, 1/2 coiled, or coiled-wavy hairs to 1 mm (or some appressed-sericeous hairs on off-coastal island plants), areoles canescent or glabrous, adaxial surface sparsely similarly hairy, glabrescent. Flowers (1–)3–8(–18) per short shoot; hypanthial tubes with coiled hairs (or some sericeous hairs on island plants). Achenes 8–11(–13) × 1.8–2.5(–2.9) mm; fruiting pedicels (1.5–)3–11(–14) mm; hypanthial tubes (6–)7–13(–16) mm; pedicel/tube ratio (20–)35–90(–120)%; fruit awns 5.3–7.5(–9.2) cm.