Plants small or medium-sized, in soft, dense mats, pale to dark green, sometimes golden or brownish, dull. Stems creeping, freely and irregularly branched to regularly pinnate; hyalodermis absent, central strand present [or absent]; pseudoparaphyllia foliose. Stem and branch leaves similar, erect-to wide-spreading, , ovate to ovate-lanceolate or lanceolate, seldom plicate; base not or scarcely decurrent; margins plane to erect or recurved, often serrulate throughout; apex short-to long-acuminate; costa double, short; alar cells differentiated, quadrate to subquadrate; laminal cells prorulose at distal, sometimes proximal, ends abaxially, rarely smooth. Specialized asexual reproduction absent. Sexual condition autoicous [rarely dioicous]; perichaetial leaves erect, triangular-lanceolate, apex gradually long-acuminate. Seta reddish. Capsule horizontal to pendent, ovoid to short-cylindric, somewhat asymmetric, ± arcuate, often constricted below mouth when dry; annulus differentiated; operculum conic, usually apiculate; peristome double; exostome teeth with external surface cross striolate proximally, coarsely papillose distally; endostome basal membrane high, segments ca. as long as teeth, keeled, cilia in groups of 1 or 2. Calyptra naked or hairs few. Spores spheric, finely papillose.
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