Plants 3-8(-13) cm, in tufts or mats. Stems dark brown or reddish brown, erect, simple or branching distally, not dendroid; rhizoids dark brown, macronemata often matted in longitudinal rows along stem, micronemata absent. Leaves green, yellowish green, reddish green, reddish brown, or black when old, contorted when dry, spreading or erect-spreading, ± flat or reflexed to arcuate-recurved when moist, orbicular, spatulate, elliptic, ovate, ovate-lanceolate, or obovate, 2-6 mm; base short-decurrent or occasionally not decurrent; margins recurved or rarely plane, green, brown, or reddish brown, 1-or 2-stratose, entire; apex acute, acuminate, obtuse, or rounded, rarely retuse, usually apiculate or rarely cuspidate, cusp usually toothed; costa percurrent or excurrent, occasionally subpercurrent or ending well before apex, distal abaxial surface smooth, abaxial stereid band present, U-shaped; medial laminal cells elongate, short-elongate, or ± isodiametric, 25-110 µm, in diagonal rows or not, not or weakly collenchymatous, walls pitted; marginal cells differentiated, short-linear or linear, in 2-4 rows. Specialized asexual reproduction absent. Sexual condition synoicous or dioicous. Seta single, yellowish, red, or reddish, 1.5-7 cm, flexuose. Capsule pendent, light or yellowish brown, yellowish green, or light green, elliptic, ovate-elliptic, or subglobose, 1.5-3 mm; operculum hemispheric; exostome greenish yellow; endostome yellow-brown, . Spores 25-70 µm.