Lianas, herbaceous. Branchlets with 4 wings and longitudinal ridges, glabrous; tendrils bifurcate. Leaves simple; stipules brownish, oval, ca. 1.5 × 1 mm, herbaceous, glabrous, apex obtuse; petiole 2-7 cm, glabrous; leaf blade oval or narrowly oval, 5-12 × 4-9 cm, glabrous, basal veins 5, lateral veins 3 or 4 pairs, veinlets conspicuously raised abaxially, base cordate or subtruncate, notch obtuse and angular, margin with 6-9 fine teeth on each side, apex cuspidate or acute. Inflorescence umbelliform, terminal or leaf-opposed; peduncle 1-2 cm, pubescent. Pedicel 2-4 mm, glabrous. Buds oval, 1.5-3 mm, apex obtuse or rounded. Calyx entire. Petals 4. Anthers oval. Lower part of ovary adnate to disk; style conical, short; stigma slightly expanded. Berry 1-1.5 × 0.8-1.4 cm, 1-or 2-seeded. Seed surface with sharp ridges, chalazal knot at lower part of seed, raphe raised, ventral holes obovate-elliptic upward to top or near middle from base. Fl. Jun-Aug, fr. Aug-Dec.