Plants coarse, glossy or dull. Leaves erect to erect-spreading, loosely imbricate when dry or moist, ovate, somewhat concave, sulcate-plicate; base ± cordate; margins in stem leaves nearly entire to denticulate distally, in branch leaves serrulate nearly to base, coarsely serrate in distal 1/3-1/2; apex acute or obtuse, sometimes short-apiculate. Sexual condition with archegoniate plants similar to antheridiate plants; perigonial and perichaetial buds distal on secondary stems near branch bases, or proximal on branches; perichaetial leaves erect, sheathing, elongate, membranous, margins entire, apex abruptly acuminate, ecostate or costa weak. Seta orange-brown to reddish. Capsule red-orange to red-brown.
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