Plants small, medium to large, mostly stiff and rigid, rarely pliant, robust to gracile, in green, yellow, olive, gray-green, brown to blackish mats. Stems creeping to ascending, repeatedly forked to sparsely fasciculately or pinnately branched, sometimes with short, tuft-like horizontal branchlets. Leaves lanceolate, ovate-or oblong-lanceolate, broadly ovate to broadly lingulate, elliptical to oblong-elliptical; margins 1-stratose to variously 2-stratose distally, mostly recurved on one or both sides, entire or erose-dentate to papillose-crenulate or crenate at apex; apices slenderly long-acuminate to rounded, rounded-obtuse to subacute, muticous or seldom with short to long awns, deeply concave to canaliculate-concave; costa single, reaching mid leaf to percurrent, entire or spurred and forked at tip, reniform to strongly flattened abaxially, 2-5-stratose in transverse section, situated in a shallow or deep, narrow-or wide-angled groove; laminal cells 1-stratose throughout to variously 2-stratose distally, elongate to linear throughout or isodiametric to shortly rectangular distally, with sinuose longitudinal walls becoming nodulose and porose with strongly incrassate walls in 2-3 rows at the base, mostly distinctly papillose with large, flat papillae covering longitudinal walls and major parts of the lumina leaving only a narrow groove in the center, occasionally more or less smooth. Perichaetial leaves differentiated, innermost elliptical to lingulate, hyaline throughout or chlorophyllose in the distal part. Seta dextrorse throughout or only with a single torsion to the right proximal to the urn and sinistrorse basally, erect, straight, smooth. Capsule brown, red-brown, or yellow-brown, straight, symmetric or nearly so, ovoid to long-cylindric; exothecial cells isodiametric to elongate, thick-walled; annulus 2-4-seriate; peristome teeth split to base or only to the middle into 2-3 terete, papillose branches. Calyptra distinctly verrucose to papillose distally. Spores spheric, finely papillose.
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