Coldenia procumbens L.

Species

Angiosperms > Boraginales > Boraginaceae > Coldenia

Characteristics

Procumbent annual herb 10–50 cm. wide or forming mats 0.5–1 m. across; stems woody at the base, ± compressed, branched and ascending, greyish-velvety.. Leaf-blades oblong, elliptic or obovate, asymmetric, 0.5–3.3 cm. long, 0.3–2.2 cm. wide, crenulate or lobulate, strigose between the nerves above with adpressed ± bulbous-based hairs pointing towards the apices of lobes making leaf appear plicate, with spreading hairs beneath; nerves in 4–6 pairs, impressed above; petiole 0.1–2 cm. long.. Flowers small, subsessile.. Calyx 2.5 mm. long, divided almost to the base into 4 lanceolate or ovate-lanceolate lobes, ± 0.6 mm. wide, often 1 lobe more developed, hairy, persistent.. Corolla white, shorter than the calyx, glabrous, soon deciduous; tube ± cylindrical, 1.5–1.7 mm. long, slightly narrowed at the throat; lobes 4, oblong to obovate, 0.3–0.5 mm. long.. Stamens 4; filaments 0.2–0.4 mm. long.. Ovary conical, 0.4–0.6 mm. long, glandular-pubescent; styles 0.1–0.2 mm. long.. Fruit 3–4 mm. long, glandular-hairy, apiculate, at first dividing into 2 then finally into 4 apiculate somewhat woody nutlets with rounded dorsal face with tubercles and irregular protuberances and angular ventral face.. Seeds flattened-ovoid, 1.8 mm. long, 1.2 mm. wide.. Fig. 11.
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Procumbent annual herb 10-50 cm wide or forming mats 0.5-1 m across; stems woody at the base, ± compressed, branched and ascending, greyish-velvety. Leaf-blades oblong, elliptic or obovate, asymmetric, 0.5-3.3 cm long, 0.3-2.2 cm wide, crenulate or lobulate, strigose between the nerves above with adpressed ± bulbous-based hairs pointing towards the apices of lobes making leaf appear plicate, with spreading hairs beneath; nerves in 4-6 pairs, impressed above; petiole 0.1-2 cm long. Flowers small, subsessile. Calyx 2.5 mm long, divided almost to the base into 4 lanceolate or ovate-lanceolate lobes, ±0.6 mm wide, often 1 lobe more developed, hairy, persistent. Corolla white, shorter than the calyx, glabrous, soon deciduous; tube ± cylindrical, 1.5-1.7 mm long, slightly narrowed at the throat; lobes 4, oblong to obovate, 0.3-0.5 mm long. Stamens 4; filaments 0.2-0.4 mm long. Ovary conical, 0.4-0.6 mm long, glandular-pubescent; styles 0.1-0.2 mm long. Fruit 3-4 mm long, glandular-hairy, apiculate, at first dividing into 2 then finally into 4 apiculate somewhat woody nutlets with rounded dorsal face with tubercles and irregular protuberances and angular ventral face. Seeds flattened-ovoid, 1.8 mm long, 1.2 mm wide.
Stems ascendingly branched, 10-50 cm long, greyish hairy. Leaves: petiole 0-5 mm long; blade 5-30 by 3-15 mm, asymmetric, oblong or obovate, crenate-dentate to lobulate, veins 4-6 on each side, impressed above, prominent beneath, strigose between veins, hairs often with bulbose base on upper, spreading on lower surface. Flowers sub-sessile. Calyx 1.5 mm long, slightly accrescent in fruit, lobes 4, lanceolate or ovate-lanceolate. Corolla white, glabrous, 1.5-1.8 mm long, tube 1-1.3 mm long, lobes ascending, 0.2-0.4 mm long, rounded. Stamens: filaments inserted above middle of corolla, included, longer than the anthers. Ovary glandular-villous; style bifid nearly to the base. Fruit 3-4 mm long, pyramidal, 4-lobed, nutlets beaked, strongly convex by vesicular, corky mesocarp dorsally, sharply angulate ventrally, with one large and one small commissural face. Fig. 3.
Herbs annual. Stems branched from base; branches prostrate or ascending, 15-40 cm, densely spreading strigose. Lower stem leaves short petiolate, oblong to obovate, 1-2 × 0.5-1.5 cm, asymmetrical, scabrous, strigose, margin coarsely crenate-lobulate; lateral veins 4-6 pairs, prominent abaxially, concave adaxially; upper leaves sessile. Flowers solitary, subsessile. Calyx 1.5-2 mm; lobes lanceolate to ovate-lanceolate, slightly enlarged in fruit, persistent, strigose. Corolla white, tubular, 2-2.5 mm, glabrous; lobes orbicular, ca. 0.3 mm. Stamens included; filaments ca. 0.3 mm, glabrous, inserted at middle of corolla tube; anthers rounded to broadly ovate, ca. 0.2 mm. Ovary fastigiate, 4-sulcate; stigma obscurely 2-cleft. Fruit broadly triangular, 3-4 mm, glandular pubescent; mericarps wrinkled, spinescent-tuberculate. Fl. Apr, fr. Jun.
Leaves with petiole 1–5(12) mm. long, on the first ones up to 40 mm. long; lamina 4–23 x 3–15 mm. (on the first leaves 25–43 x 15–32 mm.) usually oblong, sometimes circular, ovate, elliptic or obovate, plicate on the youngest leaves, with appressed hairs diverging from the nerves and converging to the teeth apex above, densely glandular-hairy below, apex rounded, base asymmetrical, cuneate on one side, obtuse to truncate on the other, margins deeply crenate to lobed.
Fruit 4–5 mm. wide, depressed-ovoid, 4-lobed, beaked and with irregular protuberances, glandular-hairy, brownish, dividing at first into 2 pairs of nutlets, later into 4 nutlets with the ventral surface angular.
Stamens inserted at c. 0.5 mm. from the base of the corolla tube; anthers c. 0.3 mm. long, suborbicular to broadly elliptic; filaments as long as the anthers, glabrous.
Ovary c. 0.4 mm. long, ovate-pyramidal, glandular-pubescent; styles slightly united at base, early concealed by the apical protuberances of the fruit.
A procumbent annual herb; stems up to 40 cm. long, radiating, angular or ± laterally compressed, much branched, densely hairy to villous.
Calyx 2.5–3.0 mm. long, a little accrescent in fruit, villous outside and inside; lobes ovate-lanceolate, subequal.
Corolla 1.5–2.2 mm. long, narrowly conical, calyptriform, early deciduous, white, glabrous.
Whitish hairy, spreading from a woody base
Small scarlet or white flowers
Flowers subsessile.
Wrinkled leaves
Life form annual
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.15 - 0.35
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

Sandy sites liable to inundation and desiccation. In plains, banks and edges of rivers and lagoons, on sandy or muddy soils that can be seasonally flooded; at elevations from 90-750 metres.
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In wet places, by muddy pools(in habit resembling Chrozophora, Forskohlea and Neurada).
Light -
Soil humidity 1-12
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 9-12

Usage

Uses medicinal
Edible -
Therapeutic use Boil (unspecified), Rheumatism (unspecified), Suppurative (unspecified), Swelling (unspecified), Tumor (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by seedlings.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Coldenia procumbens habit picture by susan brown (cc-by-sa)

Leaf

Coldenia procumbens leaf picture by Rajendra Rajendra Choure (cc-by-sa)
Coldenia procumbens leaf picture by Ramesh K (cc-by-sa)
Coldenia procumbens leaf picture by susan brown (cc-by-sa)

Flower

Coldenia procumbens flower picture by susan brown (cc-by-sa)
Coldenia procumbens flower picture by susan brown (cc-by-sa)

Distribution

Coldenia procumbens world distribution map, present in Angola, Australia, Benin, Burkina Faso, Bangladesh, Central African Republic, China, Cabo Verde, Djibouti, Ethiopia, Guinea, Guinea-Bissau, India, Iceland, Kenya, Cambodia, Lao People's Democratic Republic, Sri Lanka, Madagascar, Mali, Myanmar, Mozambique, Mauritania, Malaysia, Niger, Nigeria, Pakistan, Philippines, Papua New Guinea, Sudan, Senegal, South Sudan, Chad, Togo, Thailand, Taiwan, Province of China, Tanzania, United Republic of, Uganda, Viet Nam, South Africa, Zambia, and Zimbabwe

Conservation status

Coldenia procumbens threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:113918-1
WFO ID wfo-0000614607
COL ID WXKM
BDTFX ID -
INPN ID -
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Coldenia angolensis Lobophyllum tetrandrum Coldenia procumbens