Combretum collinum Fresen.

Species

Angiosperms > Myrtales > Combretaceae > Combretum

Characteristics

Small tree up to ± 12 m. high or a coppicing shrub; bark reddish brown or pale yellow.. Leaves opposite or alternate or verticillate; lamina coriaceous or subcoriaceous, very variable in shape, up to 22 cm. long and 8 cm. wide, upper surface drying reddish olive or yellowish brown, usually somewhat “metallic” in appearance, lower surface green, buff or silvery, glabrous (except for scales) to densely tomentose, conspicuously lepidote except when the scales are hidden by the indumentum; scales contiguous or not; lateral nerves 6–20 pairs, reticulation sometimes prominent; petiole 1–4 cm. long.. Inflorescences simple spikes or panicles up to 10 cm. long, axillary or supra-axillary from bracts or reduced leaves on the current year’s shoots (the panicle appears to be derived by the suppression of these leaves or bracts), glabrous or pubescent.. Flowers (fig. 3/10, p. 14) yellow, cream or white, fragrant.. Lower receptacle glabrous (except for scales) or hairy, lepidote; upper receptacle campanulate at the base and cupuliform at the apex, generally 3.5 mm. long, 2.5 mm. wide, or, in subsp. gazense, 5–5.5 mm. long, 3.5–4 mm. wide, glabrous (except for scales) to tomentose, lepidote.. Sepals broadly triangular.. Petals transversely elliptic to obovate or subcircular, 1.5–2.5 mm. long, 1–2.5 mm. wide, somewhat emarginate, with a claw up to 1 mm. long, glabrous (very rarely with a few hairs at the apex).. Stamen-filaments 4–4.5(–6.5 in subsp. gazense) mm. long; anthers 0.9 mm. long.. Disk 2 mm. in diameter, with pilose margin free for ± 1 mm.. Style up to 7.5 mm.. Fruit (fig. 4/10, p. 15) brown, reddish brown or greyish brown to dark purple, usually somewhat “metallic” in appearance, variable in shape, 2.5–5.9 cm. long, lepidote with the scales often contiguous, glabrous (except for scales) to densely pubescent, glossy or dull; wings variable in breadth; stipe up to 8(–15) mm. long.. Cotyledons 2, arising below soil-level on a stalk formed by their connate petioles, free part of petiole 2 mm. long (subsp. dumetorum from Angola differs in having a “normal” type of germination with the cotyledons arising above soil-level on short, quite free petioles and may be specifically distinct).. Scales (fig. 1/10, p. 12) discoid or funnel-shaped, 55–180 μ in diameter, usually extensively divided by many radial and tangential walls to give ± 10–40 marginal cells.
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A small tree. It grows 8-10 m tall. It loses many leaves during the year. The crown can be rounded or flat. The bark is reddish brown or pale brown. The leaves can be opposite or alternate. The leaf blade is 20 cm long by 8 cm wide. It varies in shape. There are 6-14 pairs of side veins. The flower clusters are 10 cm long and usually in the axils on the current year's growth. The flowers are yellow. The fruit are brown to purple and 3-5 cm long by 3-5 cm wide. They vary in shape. There are several subspecies.
Leaves opposite or alternate or verticillate; lamina up to 19(25) x 8(9) cm., very variable in shape, when dried brown or buff or golden-brown or silvery and usually somewhat “metallic” in appearance, densely tomentose to nearly glabrous (except for scales), conspicuously lepidote except when the scales are hidden by the indumentum; lateral nerves 6–14 pairs; reticulation usually (but not always) fairly prominent beneath; petiole usually up to 3(5) cm. long but very variable.
Fruit brown or greyish-brown or reddish-grey to dark-purple, usually somewhat “metallic” in appearance, 2·5–5·5 x 2–5·5 cm., variable in shape, lepidote with scales often contiguous, glabrous (except for scales) to densely hairy, glossy or dull, wings variable in breadth, stipe up to 2 cm. long.
Lower receptacle 2–4 mm. long, glabrous (except for scales) or hairy, lepidote; upper receptacle 3·5 x 2·5 mm., campanulate at the base and cupuliform at the apex, lepidote, glabrous (except for scales) to tomentose.
Petals 1·5–2·5 x 1–2·5 mm., transversely elliptic to obovate or subcircular, somewhat emarginate, unguiculate with the claw up to 1 mm. long, glabrous (rarely with a few hairs at the apex).
Scales 80–180µ in diam., extensively divided by many radial and tangential walls to give 16–40 marginal cells; scales often contiguous on the lower epidermis of the leaf.
Small semideciduous tree up to c. 17 m. high or shrub; crown rounded to flat; bark reddish-brown to grey or black or pale yellow to creamy-brown.
Inflorescences of axillary spikes or panicles up to 10 cm. long, usually in the axils of bracts or reduced leaves on the current year’s shoots.
Cotyledons 2, arising below soil level on a stalk formed by their connate petioles, free part of petiole 2 mm. long.
Disk 2 mm. in diam. with pilose margin free for c. 1 mm.
Stamen-filaments 4–4·5 mm. long; anthers 0·9 mm. long.
Flowers sessile, white or yellowish-green.
Sepals 0·8 x 1·2 mm., broadly triangular.
Style 4 mm. long, glabrous or pubescent.
Life form perennial
Growth form tree
Growth support free-standing
Foliage retention deciduous
Sexuality hermaphrodite
Pollination entomogamy
Spread -
Mature width (meter) -
Mature height (meter) 11.0
Root system -
Rooting depth (meter) 1.8
Root diameter (meter) 0.5
Flower color
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Nitrogen fixer -
Photosynthetic pathway c3

Environment

It is a tropical plant. In Namibia it grows in deep sand. It grows in wooded savanna. It grows from sea level to 1,500 m above sea level. It grows in areas with an annual rainfall of 200 mm and above. It grows in sandy and rocky soils. It can tolerate drought. It can grow in arid places.
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Savannah. Open woodlands; forest margins; deciduous thickets; and Kalahari and Lake Basin chipya woodland; occasional on anthills, in miombo woodlands; and in dry, evergreen forests; at low to medium elevations.
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Usage

The gum is edible and is eaten throughout the year.
Uses animal food bee plant charcoal environmental use food fuel gum invertebrate food material medicinal social use wood
Edible barks gums
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

It can be cut back and will re-grow.
Mode seedlings
Germination duration (days) -
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