Combretum hereroense Schinz

Russet bushwillow (en)

Species

Angiosperms > Myrtales > Combretaceae > Combretum

Characteristics

Small tree up to 8 (–12) m. high or more frequently (in E. Africa) a much branched coppicing shrub up to 5 m. high.. Bark slightly fibrous, grey-brown to blackish; branchlets reddish brown to light grey, fibrous; young shoots tomentose to densely pubescent, later becoming glabrescent, or glabrous except for the ± contiguous rufous scales.. Short horizontal spur shoots usually present.. Leaves opposite or subopposite, usually borne along the length of the spur shoots but tending to cluster towards the apex; lamina coriaceous, narrowly elliptic to broadly obovate or subcircular (especially on sucker shoots), 3–6.5(–8) cm. long, l.5–4(–4.5) cm. wide, apex slightly refuse to rounded or acute, mucronate, base cuneate to rounded, from densely tomentose to glabrous (except for the scales); scales silvery or golden and due to their wavy margins may present a rather scurfy appearance, contiguous to scattered, sometimes appearing impressed in subsp. volkensii; lateral nerves 3–9 pairs, subprominent to prominent beneath, visible above; petiole up to 5 mm. long.. Inflorescence of short, sometimes rather congested spikes up to 3 cm. long, axillary, occasionally branched, often appearing on leafless branches (especially on spur shoots) in the axils of scars of fallen leaves; peduncles ± 0.7 cm. long.. Flowers (fig. 3/20, p. 14) pale yellow to yellow, fragrant.. Lower receptacle 1.5 mm. long; upper receptacle campanulate to shortly infundibuliform, 2–3 mm. long and wide, lepidote and otherwise densely pubescent to nearly glabrous.. Sepals deltate.. Petals spathulate to very broadly obovate or subcircular, 1.5–2.5 mm. long, 1–2.5 mm. wide, emarginate, clawed, glabrous.. Stamen-filaments 2–4.5 mm. long; anthers 0.4–0.6 mm. long.. Disk with a pilose margin free for 0.7–0.8 mm.. Style 3–4 mm. long.. Fruit (fig. 5/20, p. 16) oval to subcircular in outline, up to 3.5 cm. long and wide, often much smaller, glabrous except for the scales; apical peg absent or very short; stipe up to 11 mm. long.. Cotyledons with petioles 10–11 mm. long; germination epigeal.. Scales (fig. 2/20, p. 13) irregularly undulate in outline, (50–)75–100 μ in diameter, marginal cells 8–12, mostly retusely scalloped.
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Leaves opposite or subopposite, often borne on short shoots; lamina usually c. 2–7 x 1–4·5 cm., from very narrowly elliptic to very broadly obovate or subcircular (especially on sucker shoots when they may be as much as 5·5 cm. broad and broader than long), from densely tomentose to nearly glabrous (except for scales), densely golden-(rarely silvery-)lepidote beneath, scales usually contiguous but sometimes hidden by the indumentum, apex sometimes acute but often rounded or even retuse, base cuneate to rounded; lateral nerves usually only 3–4(5) pairs, prominent beneath; petiole up to 5 mm. long.
A small tree. It can be up to 11 m tall, but is often 3-5 m tall. The trunk is often crooked and can be 23 cm across. The leaves can be oval or heart shaped. The leaves are 1.9-5 cm long by 1.3-2.5 cm wide. The are dark green and shiny above and yellow-green underneath. Often there are dense velvety hairs underneath. The flowers are small and white or greenish-yellow. They occur in long spikes. The fruit are 2.5 cm long by 1.9 cm wide. They have 4 wings. They are bright red.
Inflorescences of short rather congested spikes up to c. 3 cm. long, occasionally branched, axillary, often appearing on leafless branchlets in the axils of the scars of the fallen leaves; rhachis tomentose to glabrous (except for scales); bracts 3 mm. long, filiform, often somewhat persistent.
Small tree up to 9(12) m. high or more rarely a shrub, often with arching stems, frequently flowering before the leaves; crown dense, irregular; bark creamy-grey to grey-black, fissured; branchlets usually tomentose to densely pubescent when young but often quickly glabrescent.
Fruit usually c. 2 x 2 cm. but sometimes smaller and occasionally up to 3·5 x 3·5 cm., subcircular in outline, densely reddish-or golden-lepidote, apical peg absent or very short, wings c. 5(10) mm. wide, subcoriaceous, stipe up to 11 mm. long.
Scales irregularly undulate in outline, marginal cells 8–12 mostly retusely scalloped, delimited by usually 8 radial walls, 8 tangential walls and sometimes a few extra radial walls.
Lower receptacle 2·3–3 mm. long; upper receptacle 2·5–3 x 2–3 mm., campanulate or shortly infundibuliform, lepidote and otherwise pubescent to nearly glabrous.
Petals 1·5–2·5 x 1–2·5 mm., broadly to very broadly obovate or subcircular or very broadly transversely elliptic, emarginate, shortly unguiculate, glabrous.
Small tree or shrub, up to 9 m high. Fruit 20 mm long, russet-coloured. Flowers yellow; August to January. Tree or shrub savanna.
Cotyledons 2 1·5 x 2·5 cm., transversely elliptic, arising above soil level; petioles 10–11 mm. long.
Disk c. 2 mm. in diam. with pilose margin free for 0·7–0·8 mm.
Stamen-filaments 3–1·5 mm. long; anthers 0·4–0·6 mm. long.
Style 3·5–4·5 mm. long, not expanded at the apex.
Sepals 1–1·5 x l–1·5 mm., deltate.
Flowers sessile, yellow.
Life form perennial
Growth form
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 9.0 - 10.0
Root system -
Rooting depth (meter) 1.4
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

A tropical plant. It often grows in stony and shallow soils. It grows at medium to low altitudes in open wooded grassland. It can grow in salty soils. It can tolerate drought. It grows between 30-1,465 m above sea level. It can grow in arid places.
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Mixed open or savannah woodland at elevations from 50-2,700 metres.
Light 4-6
Soil humidity 3-5
Soil texture 3-4
Soil acidity -
Soil nutriment -
Hardiness (USDA) 10-10

Usage

The dried fruit are used as a tea substitute. The seeds are considered poisonous and are removed. The fruit are also eaten fresh. The leaves are dried and used for tea.
Uses animal food environmental use food fuel material medicinal poison social use tea
Edible fruits gums leaves
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Plants can be grown from seeds. The seeds are best removed from the outer covering and soaked in water for a few hours. Seeds germinate in 1-2 weeks. Seedlings can be transplanted at the 3 leaf stage. It can be cut back and will re-grow.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) 1
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Leaf

Combretum hereroense leaf picture by Lusajo Mwakibinga (cc-by-sa)

Distribution

Combretum hereroense world distribution map, present in South Africa

Conservation status

Combretum hereroense threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:170145-1
WFO ID wfo-0000616365
COL ID 5ZRCS
BDTFX ID -
INPN ID -
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Combretum hereroense Combretum borumense Combretum rhodesicum Combretum sambesiacum Combretum usaramense Combretum villosissimum Combretum porphyrolepis Combretum rautanenii Combretum eilkerianum Combretum transvaalense

Lower taxons

Combretum hereroense subsp. grotei Combretum hereroense var. parvifolium Combretum hereroense var. villosissimum Combretum hereroense subsp. volkensii