Shrubs or small trees, stellate-pubescent. Leaves opposite, subsessile to petiolate, simple, ±flat, entire or irregularly undulate-dentate. Flowers 1–5 (–7) and terminal to short branchlets or solitary and axillary, bisexual, 4-merous. Pedicels bibracteolate. Calyx cup-shaped, entire or 4 (or 4 + 4)-lobed. Corolla mostly sympetalous, mostly tubular, stellate-hairy; tips of lobes slightly inflexed. Stamens 4 + 4, inserted at base of disc; filaments linear, variably broadened at base, glabrous, antipetalous ones shorter; anthers included or exserted, versatile, oblong to lanceolate, introrse, not apiculate, abaxially with reddish brown connective. Disc 8-lobed, glabrous. Carpels 4, united at base, variably tomentose; style filiform, attached subapically to adaxial margin of ovaries; stigma minutely 4-lobed. Fruit of 4 cocci.