Plants 4.5-15 mm, in dense cushions to loose mats, olivaceous to dark black-green. Leaves oval to ovate-lanceolate, keeled distally, margins plane, incurved, or recurved, distal lamina 1-or 2-stratose, specialized laminal and marginal chlorophyllose structures absent, usually long-awned; basal cells usually rectangular with straight walls; mid leaf and distal cells quadrate to short rectangular, straight or sinuose, usually thick-walled. Gemmae absent. Sexual condition autoicous or dioicous; perichaetial leaves enlarged. Seta short to long, straight. Capsule erect, immersed to exserted, symmetric, campanulate, ovoid to cylindrical, rarely cupulate; annulus poorly differentiated; operculum conic to rostrate, falling detached from the columella. Calyptra campanulate, becoming cucullate with age, erose at base, large, covering 1/2 to all of capsule, plicate.
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