Plants somewhat stiff. Stems with shoot and branch apices often pale; paraphyllia sometimes present only in youngest shoot portions; axillary hairs few, weak. Stem leaves gradually or suddenly narrowed to apex, plane or slightly concave; alar cells usually many, region large, usually strongly widened marginally, reaching from margin to costa or almost so; medial laminal cells 12-55 × 4-11 µm. Branch leaves usually narrower, more strongly falcate. Perichaetia with inner leaves gradually or suddenly narrowed to apex, plicate, margins denticulate or strongly so distally, apex acuminate, laminal cells smooth.