Plants small to medium-sized, seldom large, in loose mats, light green, dull. Stems usually creeping, pinnate or subpinnate; hyalodermis absent, central strand present; pseudoparaphyllia triangular to ovate-lanceolate. Stem and branch leaves differentiated. Stem leaves spreading, , ovate-lanceolate to triangular-acuminate, not or weakly plicate; base broadly decurrent; margins usually recurved basally, plane distally, denticulate; apex acuminate from near base or mid leaf; costa usually double, short or to 1/4 leaf length; alar cells weakly differentiated, short-to long-rectangular; laminal cells distally prorulose abaxially. Branch leaves longer lanceolate, smaller. Specialized asexual reproduction absent. Sexual condition dioicous; outer perichaetial leaves weakly reflexed, inner leaves erect, ovate-lanceolate, apex acuminate. Seta reddish, smooth, occasionally scabrous near capsule. Capsule nodding to horizontal, short-elliptic to cylindric and curved, not or somewhat constricted below mouth; annulus somewhat differentiated, 1-or 2-seriate, cells large; operculum conic-apiculate; peristome double; exostome teeth with external surface with zigzag central line, lamellose, striolate proximally, papillose distally; endostome basal membrane high, segments as high as exostome, keeled, cilia 1-3, elongate. Calyptra weakly prorulose distally, hairy. Spores spheric, finely papillose.
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