Trees deciduous, monoecious. Branchlets with chambered pith. Terminal buds oblong, naked. Leaves odd-pinnate; leaflets (5 or)7 or 9(or 11), margin serrate. Inflorescences lateral or terminal on old or new growth, pendulous; male and female inflorescences separate: male spikes in clusters of 3-5, lateral on old growth; female spike solitary, terminal on new growth. Flowers anemophilous. Male flowers with an entire bract; bracteoles 2; sepals 2; stamens 20-31, anthers pubescent. Female flowers subtended by a small, entire bract, adnate to bracteoles and virtually submerged in wing complex; bracteoles 2, united and adnate to ovary; sepals 4, adnate to ovary, free at apex; style short; stigmas commissural, 2-lobed, plumose. Fruiting spike elongate, pendulous. Fruit a disc-winged nutlet, 2-4-chambered at base. Germination most likely hypogeal.
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