Cyrilla racemiflora L.

Swamp titi (en)

Species

Angiosperms > Ericales > Cyrillaceae > Cyrilla

Characteristics

Shrub or small tree, up to 7-15 m tall, sometimes multistemmed, occasionally spreading with vine-like branches, rarely branched at base. Leaf scars prominent. Stipules inconspicuous (caducous) or absent; petiole usually short (nearly subsessile), slightly winged, 0.4-1.3 cm long; blade subcoriaceous to coriaceous, oblanceolate to elliptic, occasionally linear, 2-11(-12) (including petiole) x 0.4-2.5(-3) cm, margin usually slightly reflexed, apex acute to rounded, sometimes emarginated (rarely apiculate), base narrowly cuneate to attenuate; midvein generally depressed above, prominent beneath, densely reticulate, slightly prominent or obscure above, secondary veins straight to slightly arcuate, divergent, irregularly anastomosing. Inflorescence racemose, axillary, usually near end of branches, glabrous; peduncle 0.4-1.7 cm long (before the first bract); rachis up to 14 cm long, usually somewhat ridged; often numerous flowers, erect or pendulous; bract lanceolate-subulate, 0.8-2.4(-3.5) mm long, acuminate to very acute, basally thickened and usually costate, persistent; bracteoles 2, usually above middle of pedicel, alternate or opposite, lanceolate, 1-1.5(-1.6) mm long, persistent; pedicels ascending or slightly patent, alternate or subopposite, 1.1-2 mm long (up to 4 mm when fruiting), usually pink. Flowers actinomorphic; sepals free, or basally connate, 5, pink, coriaceous, usually with basal thickening, lanceolate-ovate to deltoid, 0.6-1.5(-1.6) x up to 0.7(-1) mm, apex acute to acuminate, glabrous, persistent; petals imbricate or slightly contorted, free or shortly fused, 5, white, usually more membranaceous along margin, oblong to oblong-lanceolate or oblong-elliptic, 2(-2.4) x up to 1.1 mm, margins sometimes inrolled (giving the appearance of long acuminate), apex acute, obtuse or rounded, glabrous to rarely glandular on lower inner surface; stamens 5, about 2/3 as long as petals (always shorter), filaments flattened, 1.1-1.4 mm long, anthers terete, linear-subulate, up to 0.4 mm broad, very occasionally with mucronate tip, versatile, dehiscing longitudinally, pink or purple; nectary disc sometimes present; ovary 2-3(rarely-4)-locular, oblong to slightly ovate, wrinkled when dried, glabrous, up to 1.6(-2) mm long, ovules usually 2, pendulous from apex of locule, style 1, short, stout, stigma entire, or 2-3-lobed, corresponding to number of locules. Fruit dry, indehiscent, green immature, red (pink) at maturity, subglobose to globose or conic-ellipsoid, 2.5-(3) x up to (0.9-)2(-3) mm, usually smooth (rarely slightly rugulose), glabrous, stigma usually persistent; seeds 1 per locule, elongate.
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Trees (in Panama) with thick, rough, brown bark; diam ca 45 cm; branches glabrous, terete; branchlets subterete to angulate, glabrous. Leaves crowded towards the ends of the branches, the blade oblanceolate, narrowly cuneate at base, acute to subobtuse at apex, recurved marginally, coriaceous, glabrous, the midvein prom-inent beneath, densely reticulate, 2-9 cm long, 1-2.4 cm broad; petiole to! 4 mm long, glabrous. Racemes 3.5-1.0 cm long, borne in the upper axils, erect to pendu-lous, glabrous; peduncles 4-10 mm long, glabrous; bracts 0.5-1.5 mm long, lance-olate, glabrous, persistent, ascending in bud, spreading to descending after anthesis. Flowers ascending in bud, spreading at anthesis, pedicellate, the pedicel 1-2 mm long, glabrous; bracteoles paired, alternate to opposite, 0.5-1 mm long, lanceolate, basally thickened, usually persistent, borne above the middle of the pedicel, spread- ing after anthesis; sepals 5, free, ovate-lanceolate to deltoid, acute at apex, 0.4-0.7 mm long, imbricate in bud; petals 5, inserted on a small glandular disk, o0blong-lanceolate to oblong-elliptic, acute at apex; stamens opposite the petals, 1.5-2 mm long, the filaments terete, the anthers versatile; ovaries sessile, 2-3(-4)-locular, the style persistent, the stigma 2-4-lobed. Fruits dry, globose, ovoid to oblong-ovoid, 1.6-2.7 mm in diam; seeds elongate; cotyledons short, inferior; radicle superior.
Shrub or small tree to 10(–20) m; lvs leathery, persistent through much of the winter, oblanceolate to obovate, 5–10 cm, the petiole 5–8 mm; racemes several, borne just below the leafy branches of the season, spreading, slender, 6–15 cm, floriferous to the base; bracts subulate, equaling or longer than the pedicels; pet white, oblanceolate, 3 mm, slightly surpassing the stamens; fr gray, round-ovoid to depressed-globose, 2–3 mm, excluding the persistent short, 2–3-lobed style; 2n=40. Swamps and wet woods on the coastal plain; se. Va. to Tex., s. into n. S. Amer. June.
Plants to 10 m, frequently forming dense thickets. Leaves: petiole distinct; blade green, oblanceolate to elliptic, 1-10 × 0.5-2.5 cm, chartaceous to coriaceous, base cuneate, apex acute, rounded, or emarginate. Flowers fragrant; petals white or creamy white, 2.5-3.5 mm. Fruits ovoid to subglobose, 2-2.5 mm.
Life form perennial
Growth form shrub
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
Pollination entomogamy
Spread -
Mature width (meter) 2.0
Mature height (meter) 6.0 - 9.0
Root system -
Rooting depth (meter) 0.6
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway c3

Environment

Rich shaded river bottoms, the borders of sandy swamps and shallow ponds of the coastal pine-belt. Also found on high, sandy, exposed ridges rising above streams.
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Rich shaded river bottoms, the borders of sandy swamps and shallow ponds of the coastal pine-belt. Also found on high, sandy, exposed ridges rising above streams.
Light 3-7
Soil humidity 5-7
Soil texture 2-5
Soil acidity 2-6
Soil nutriment -
Hardiness (USDA) 6-8

Usage

Uses: Planted as an ornamental in the USA. In the West Indies, the species is esteemed among honey producers due to its nectar-rich flowers, which are present over an extended period of time (Ståhl, 2004).
Uses charcoal environmental use invertebrate food material medicinal ornamental seasoning timber wood
Edible -
Therapeutic use Astringent (unspecified), Hemorrhage (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by cuttings or seedlings.
Mode cuttings seedlings
Germination duration (days) -
Germination temperacture (C°) 21
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -23
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Leaf

Cyrilla racemiflora leaf picture by Luke Luke (cc-by-sa)
Cyrilla racemiflora leaf picture by Christa Chasteen (cc-by-sa)
Cyrilla racemiflora leaf picture by Paris B (cc-by-sa)

Flower

Cyrilla racemiflora flower picture by Luke Luke (cc-by-sa)
Cyrilla racemiflora flower picture by Eli Small (cc-by-sa)
Cyrilla racemiflora flower picture by Mary Olsen (cc-by-sa)

Fruit

Cyrilla racemiflora fruit picture by Luke Luke (cc-by-sa)
Cyrilla racemiflora fruit picture by Eli Small (cc-by-sa)
Cyrilla racemiflora fruit picture by John Cumper (cc-by-sa)

Distribution

Cyrilla racemiflora world distribution map, present in Belize, Brazil, Colombia, Cuba, Dominica, Dominican Republic, Guadeloupe, Guatemala, Guyana, Honduras, Haiti, Jamaica, Mexico, Nicaragua, Panama, Puerto Rico, United States of America, and Venezuela (Bolivarian Republic of)

Conservation status

Cyrilla racemiflora threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:315951-1
WFO ID wfo-0000634813
COL ID 33JMD
BDTFX ID -
INPN ID 629522
Wikipedia (EN)
Wikipedia (FR) Link

Synonyms

Andromeda plumata Cyrilla antillana Cyrilla arida Cyrilla brevifolia Cyrilla caroliniana Cyrilla perrottetii Stachyanthemum schomburgkii Itea floribunda Itea caroliniana Itea caroliana Itea cyrilla Itea cyrilla l'hér. Cyrilla fuscata Cyrilla parvifolia Cyrilla parvifolia Cyrilla polystachia Cyrilla racemiflora var. brevifolia Cyrilla racemiflora subsp. parvifolia Cyrilla racemiflora var. parvifolia Cyrilla racemiflora var. subglobosa Cyrilla racemiflora var. racemiflora Cyrilla racemiflora var. parvifolia Cyrilla antillana var. perrottetii Cyrilla racemifolia var. parvifolia Cyrilla racemiflora