Plants small. Stems creeping, 1-or 2-pinnate; paraphyllia filamentous, unbranched; axillary hairs 2-celled. Stem leaves broadly triangular to broadly ovate; margins recurved at least basally, crenulate-papillose; costa 3/4 leaf length to subpercurrent; laminal cells ± quadrate, multipapillose [1-papillose] on both surfaces. Branch leaves ovate to oblong-ovate; margins plane, crenulate-papillose; apex obtuse to short-acuminate; costa 1/2-7/8 leaf length, covered with cells apically or not; laminal cells ± quadrate, multipapillose [1-papillose] on both surfaces. Sexual condition autoicous; perichaetial leaves oblong-lanceolate to oblong-ovate, margins subentire to ciliate, costa percurrent to excurrent, laminal cells papillose. Seta smooth or papillose. Capsule inclined to horizontal, cylindric, asymmetric; annulus 2-or 3-seriate; operculum obliquely long-rostrate; endostome segments not or narrowly perforate, cilia 1-3.