Rhizome 3-15 mm in diam. (without scales), not white waxy. Scales red-brown or nearly black, with pale border from base to apex or without pale border, narrowed evenly toward apex or flat and nearly acicular, narrowed abruptly from a broad base, often curling backward, peltate, 4-8 × 0.5-1.5 mm, without multiseptate hairs, toothed. Stipe pale, adaxially grooved, 4-50 cm, glabrous or with few scales; lamina compound, bipinnate or quadripinnate toward base and in middle part, deltoid and broadest toward base, 16-90 × 13-50 cm, glabrous, not or slightly dimorphic. Longest petiolules 4-35 mm; pinnae deltoid; longest pinnae 8-45 × 5-30 cm; pinnules of at least larger pinnae anadromous, deltoid; longest pinnules 70-200 × 40-110 mm; ultimate pinnae linear-oblong or narrowly ovate, lobed almost to midrib or only shallowly lobed; ultimate segments 5-27 × 2-6 mm. Rachises and costae glabrous. Margins of each pinna not thickened. Veins in sterile ultimate lobes pinnate (or forked in very narrow lobes), reaching margin; false veins present. Sori separate, several per segment, at forking point of veins; indusium also attached along sides, pouch-shaped, oblong, longer than wide or ± as wide as long, 1-1.3 × 0.5-1 mm, upper margin not elongated, truncate or slightly rounded, separated from or even with lamina margin; lamina generally extending into a tooth at both sides of a sorus.
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