Rhizome bearing only scales. Roots restricted to the ventral side of the lateral buds. Scales yellowish, light brown, or nearly black, with pale border extending from base to apex, quickly diminishing or disappearing into the apex, or without pale border, not bearing multicellular hairs, lacking marginal setae (or with marginal setae in D. novoguineense), entire, smooth on adaxial surface, peltate, or basifixed with a cordate base with greatly overlapping lobes (in D. viscidulumand D. urceolatum). Stipe articulated on phyllopodia, grooved, bearing hairs and/or scales when young. Lamina compound, pinnate with strongly dissected pinnae or bipinnate towards the base and in the middle part, elongate and narrowed towards the base, bearing multicellular hairs, not or slightly dimorphous. Hairs between veins on each surface present or not. Pinnae linear-triangular. Pinnulae or pinna lobes linear oblong. Ultimate segments obtuse, without a dominant tooth. Pinnules of at least the larger pinnae catadromous. Rachis adaxially raised. Leaf axes, at least rachises, hairy. Veins in ultimate lobes simple or forked, not reaching the margin. False veins wanting. Sori indusiate, separate, frequently single on a segment, facing midveins at the forking point of veins or (rarely) at the bending point. Indusium scaly, attached at the narrow, cordate base only, or attached at the base and only part way up at the sides, or also attached along the sides, pouch-shaped, or attached at the broad base and hardly or not at the sides, or very small, inconspicuous.
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Fronds more or less copiously persistently hairy; lamina thin; Indusium small, of various shapes.