Plants summer-green. Rhizome slender, creeping, dark brown, 2-3 mm in diam., with sparse yellow-brown, lanceolate, membranous scales; fronds distant. Lamina subdimorphic, stipe of fertile frond prominently longer than in sterile frond; fertile frond up to 90 cm; stipe dark brown at base, upward stramin-eous, 18-45 cm, 1-3 mm in diam. at base, base sparsely scaly, scales similar to those on rhizome, upward with more brown to dark brown, shiny lanceolate scales and yellow-brown small nodose hairs, then scales and hairs gradually deciduous; lamina narrowly deltoid, base not narrowed, or ovate, 16-45 cm, base slightly narrowed, apex acuminate to long acuminate, sometimes abruptly narrowed; free pinnae 7-12 pairs, mostly alternate, spreading or slightly ascending, sessile, base truncate or broadly cuneate, occasionally shallowly cordate, pinnatipartite, apex lanceolate or oblong-lanceolate, rarely subacute; upper pinnae symmetrical or nearly symmetrical, basal pinnae asymmetrical with basiscopic lobes longer; pinna lobes 7-17 pairs, contiguous, ascending or subspreading, ligulate-oblong, margin toothed or repand, rarely pinnatipartite or subentire, apex rounded; middle pinna lobes 1-2 cm × 5-8 mm; veins pinnate with 1-8 pairs of veinlets, visible on both sides, forked or simple. Lamina herbaceous, green or pale green when dry, darker on adaxial side; both sides of rachis, costae, and veins with sparse gray-white, pale brown nodose soft hairs, hairs on abaxial side of costae thick, long. Sori shortly linear, straight or slightly oblique, 1-7 pairs per pinna lobe, mostly single, medial or inframedial, proximate end of sori near costa, double on basal acroscopic veinlets; indusia light brown when young, yellow-brown when old, membranous, hairy or glabrous, margin flat, lacerate, then erose, long or shortly ciliate. Spores orbicular in polar view, semicircular in equatorial view, perispore prominent, hyaline, surface ornamentation with mostly clavate, apically truncate, sometimes narrow flakelike projections.
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