Plants green to dark green, usually with a reddish cast. Stems to 1 cm, central strand present. Stem leaves spirally twisted and appressed to weakly spreading when dry, weakly spreading and not keeled when moist, monomorphic, ovate to ovate-lanceolate, grooved adaxially along the costa, especially near leaf apex, to 0.6-1 mm, base ovate in shape, margins broadly recurved to revolute to apex of leaf, entire, apex conic-acute to obtuse, not fragile, commonly ending in a conical cell; costa percurrent, not strongly spurred, widened medially, with a broad adaxial 1-stratose pad of thin-walled cells, adaxial costal cells quadrate to short-rectangular, 7-9 cells wide at mid leaf, guide cells in 2(-3) layers; basal laminal cells differentiated medially or across the leaf, walls thin to evenly thickened, short-rectangular, not perforated; distal laminal cells 8-10 µm wide, 1:1 or occasionally 1:2, nearly smooth or papillae often as a thickened and irregularly shaped lens covering the lumen, lumens quadrate, walls thin or weakly thickened, weakly convex on both sides of lamina, distal leaf margins 1-stratose. Specialized asexual reproduction by rhizoidal tubers or axillary gemmae of (2-)4-6(-10) cells. Sexual condition only perichaetiate plants seen. Sporophytes unknown. Distal laminal KOH reaction blotchy or irregularly blushing red at mid leaf or above, otherwise colorless or light yellow.