Didymodon tophaceus (Brid.) Lisa

Didymodon moss (en)

Species

Bryophytes > Pottiales > Pottiaceae > Didymodon

Characteristics

Plants small to medium, in dense cushions, light green above, light brown below; terricolous or saxicolous. Stems 10-50 mm tall, frequently branched, especially in taller plants; in section round, central strand small, inner cortical cells large, thin-walled, outer cortical cells in 2-3 rows, smaller, incrassate, yellowish, outermost row sub-stereids. Leaves appressed to slightly contorted dry, patent wet, shape extremely variable on same plant, ovate-acuminate to lanceolate or ligulate, 1.0-1.8 mm long; apex acute to broadly rounded; margins entire, reflexed to recurved below, plane to reflexed above. Costa percurrent to ending 10 cells below rounded apices, ventral and dorsal superficial cells quadrate to short-rectangular, incrassate, smooth; in section round, weakly differentiated, guide cells 2, ventral cells in 1-2 layers, incrassate, dorsal stereid band 1-2 rows thick, frequently cells substereids only, dorsal surface cells larger, similar to ventral cells, incrassate. Upper laminal cells quadrate to rounded-quadrate, incrassate, frequently with thickened corners in upper leaf, some cells with low, blunt papillae frequently obvious only in leaf sections; basal cells rectangular to short-rectangular, thin-walled, smooth. Perichaetia terminal, becoming lateral through innovation, leaves ligulate, 1.5 mm long; apex obtuse to broadly rounded; leaf cells occasionally somewhat larger than vegetative leaf cells. Seta erect, 7-8 mm long, reddish; capsule short-cylindrical, 1.5 mm long, reddish to yellowish red; peristome absent; operculum rostrate, 0.4 mm long; spores round, 12-15 µm, weakly granulate.
More
Plants green to olive green or tan. Stems to 1.5 cm, central strand present. Stem leaves weakly spreading or rarely appressed when dry, spreading and often keeled when moist, monomorphic, ovate to long-elliptic but occasionally ovate-lanceolate, adaxially grooved along costa or broadly channeled, 1-2 mm, base ovate, margins recurved in proximal 3/4 of leaf and often decurrent on stem, entire, apex broadly acute to obtuse or rounded, often in same collection, not fragile; costa percurrent or ending 1-6 cells below the apex, tapering or rarely spurred above, sometimes much broadened at the base, pad of cells absent, adaxial costal cells elongate at least in the central 2 rows, 4 cells wide at mid leaf, guide cells in 1 layer; basal laminal cells weakly differentiated medially, quadrate or short-rectangular, walls usually thickened; distal laminal cells 9-12(-14) µm wide, 1:1, papillae usually absent, rarely simple and 1-2 per lumen, lumens rounded-quadrate or rounded, walls thin to thickened, weakly convex on both sides, 1-stratose. Specialized asexual reproduction specialized structures absent. Seta 0.7-1.2 cm. Capsule 0.6-1.5 mm; peristome teeth absent, rudimentary or short and 16, ligulate, cleft to near base, untwisted, to 220 µm. Spores 12-14(-20) µm. Distal laminal KOH reaction red-to orange-brown, occasionally yellow-brown.
Plants in dense cushions. Stems 10-50 mm long, central strand small. Leaves ovate-acuminate to lanceolate or ligulate, 1.0-1.8 mm long; apex acute to rounded; margins reflexed to recurved below; costa percurrent to ending below, ventral stereid band absent. Upper laminal cells rounded-quadrate, incrassate, some cells with low papillae. Capsule short-cylindrical, peristome absent.
Life form -
Growth form -
Growth support -
Foliage retention deciduous
Sexuality -
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.01 - 0.05
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Light -
Soil humidity 1-12
Soil texture 7-8
Soil acidity -
Soil nutriment -
Hardiness (USDA) -

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Distribution

Didymodon tophaceus world distribution map, present in South Africa

Identifiers

LSID 35132203
WFO ID wfo-0001164498
COL ID 35YWT
BDTFX ID -
INPN ID 5300
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Dactylhymenium pringlei Barbula wildii Weissia amblyphylla Didymodon ceratodonteus Didymodon tophaceus Pottia ceratodontea Gymnostomum ceratodonteum Gymnostomum platyphyllum Husnotiella pringlei Hymenostylium platyphyllum Trichostomum tophaceum Desmatodon hendersonii Barbula pringlei Barbula tophacea Didymodon diversifolius Didymodon hendersonii Didymodon ligulifolius Didymodon planotophaceus Trichostomum limosum Didymodon bosniacus Didymodon spadiceus var. siluricus Didymodon tophaceus var. breidleri Didymodon trifarius var. tophaceus Didymodon tophaceus var. ligulifolius

Lower taxons

Didymodon tophaceus var. elatus Didymodon tophaceus var. excurrens Didymodon tophaceus var. laxus Didymodon tophaceus var. linearis