Didymodon umbrosus (Müll.Hal.) R.H.Zander

Species

Bryophytes > Pottiales > Pottiaceae > Didymodon

Characteristics

Plants usually green to very bright green distally, contrasting with light tan proximally. Stems to 1(-1.5) cm, central strand present. Stem leaves spreading-incurved and twisted to incurved-appressed when dry, spreading to spreading-recurved and not keeled when moist, monomorphic, long-lanceolate, broadly concave adaxially across leaf, usually 1-2.5(-4) mm, base sharply differentiated in shape, ovate, margins usually plane, entire, apex narrowly acute, not fragile; costa percurrent or ending a few cells below the apex, broader at mid leaf, occasionally weakly spurred, with a low adaxial pad of cells, adaxial costal cells rectangular, 4-6 cells wide at mid leaf, guide cells in 1-2 layers; basal laminal cells strongly differentiated medially, rectangular, walls very thin or hyaline, often perforated by transverse slits; distal laminal cells in rather distinct rows, 7-12 wide, 1:1 or occasionally longitudinally elongate, papillae usually absent, lumens oval to rounded-quadrate or rounded-rectangular, walls convex on both sides of lamina, 2-stratose in one or more rows along margins. Specialized asexual reproduction by multicellular tubers on proximal rhizoids. Seta 0.7-1 cm. Capsule 1-1.9 mm; peristome teeth 32, linear, weakly twisted, to 600 µm. Spores 11-15 µm. Distal laminal KOH reaction variously negative or yellow-or orange-or red-brown.
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Plants small, caespitose, green to yellow-green; terricolous. Stems to 10 mm high, simple; in section round, central strand very small, inner cortical cells lax, outer cortical cells in 1-2 rows, small, incrassate, reddish. Leaves incurled dry, erect wet; lanceolate to ovate-acuminate, 1.5-2.0 mm long, lamina unistratose; apex broadly acute; margins bistratose, cells slightly smaller than laminal cells. Costa percurrent, ventral superficial cells quadrate, smooth, dorsal superficial cells rectangular, smooth; in section oval, guide cells 4-5, ventral cells in 1(-2) layer(s), large, mammillose, dorsal stereid band 2-3 cells thick, dorsal surface cells undifferentiated. Upper laminal cells irregularly quadrate to short-rectangular, smooth or nearly so; marginal cells quadrate, smaller; basal cells rectangular, thin-walled; basal marginal cells in 1-3 rows, narrowly rectangular. Dioicous. Perichaetial leaves undifferentiated. Seta 8-10 mm long, reddish yellow; capsule short-cylindrical, 1.5 mm long; peristome divided to base, filaments erect, papillose, yellowish red; operculum rostrate, 0.8 mm long; spores sub round, 15 µm, granulate.
Plants caespitose. Stems up to 10 mm long, central strand very small. Leaves incurled when dry, lanceolate to ovate-lanceolate, 1.5-2.0 mm long; apex broadly acute; margins bistratose; costa percurrent, ventral stereid band absent. Upper laminal cells irregularly quadrate to short-rectangular, ± smooth, basal cells rectangular, thin-walled. Capsule short-cylindrical, peristome teeth divided.
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Foliage retention deciduous
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Mature height (meter) 0.01
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Cultivation

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Distribution

Didymodon umbrosus world distribution map, present in South Africa

Identifiers

LSID 35140559
WFO ID wfo-0001169397
COL ID 35YWW
BDTFX ID -
INPN ID 434202
Wikipedia (EN)
Wikipedia (FR)

Synonyms

Trichostomopsis crispifolia Didymodon umbrosus Asteriscium flavisetum Barbula flaviseta Barbula umbrosa Trichostomopsis umbrosa Didymodon australasiae var. umbrosus