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Male flowers with 6–8 sepals in 2 whorls; petals absent; stamens 3–6, fused into a cylindric or subglobular, sometimes flattened, sessile or stipitate synandrium; anther-thecae oblong, parallel, dehiscing longitudinally.
Drupelets subovoid, topped by the remains of the style and stigma; exocarp smooth and sometimes shining; endocarp crustaceous, verrucose or covered on the upper side with short prickles much enlarged at the base.
Female flowers with 6 sepals; petals absent; carpels 3–6, with thickened, recurved stigmas.
Herbaceous twiners: Leaves simple, with long petioles, entire or lobed.
Male inflorescences of axillary, long-pedunculate racemes.
Female inflorescences similar to the male.
Seeds with thick fleshy endosperm.
