Rhizome shortly erect or decumbent, glabrous, or apex with sparse, dark brown scales; fronds caespitose; stipe brown at base, upper part and rachis light brown, densely hairy, hairs shiny, curly long nodose, shallowly grooved on adaxial side, rounded abaxially; lamina elliptic or narrowly elliptic, base slightly narrowed; pinnae approximate, alternate or opposite, sickle-shaped, lanceolate, base sessile, asymmetrical, acroscopic base deltoid-auriculate, or rounded-obtuse, basiscopic base rounded, margin entire or repand, sometimes serrate, apex acuminate, acute, rounded, or obtuse; supramedial pinnae spreading, lower pinnae reflexed, shortened, basal pinnae sometimes ca. 1/2 of middle pinnae, occasionally not reduced; veins prominent on both surfaces of lamina, midrib densely hairy on abaxial surface of lamina, hairs nodose, sparsely hairy or glabrous on abaxial surface. Sori and indusia mostly long linear, up to more than 2/3 or on entire length of veinlet, or shorter; indusial surface ± with long nodose hairs.
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