Plants small to large. Stems pinnate or irregularly pinnate. Stem leaves falcate or sometimes straight, ovate-lanceolate, ovate, triangular-ovate, rounded-triangular, or broadly ovate, gradually narrowed to apex, concave, 0.9-5.1 × 0.4-1.6 mm; base erect to erectopatent, insertion slightly curved; margins entire or occasionally very finely denticulate; apex [obtuse] acute, or short-to long-acuminate, acumen gradually differentiated, in straight-leafed plants plane or at most weakly furrowed; costa single to mid leaf or beyond, ending well before apex; alar region transversely triangular, reaching from margin to costa or almost so; ratio of medial laminal cell length (µm) to leaf length (mm) 17.9-35.5 (very rarely to 38.2). Sexual condition dioicous.
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