Plants medium-sized to large. Stems irregularly and often sparsely branched. Stem leaves patent, spreading, or squarrose, ovate, broadly ovate, or triangular-ovate, suddenly or gradually narrowed to apex, concave or strongly concave, (1-)1.7-3.6 × 0.6-1.5 mm; base erect, subclasping, insertion usually deeply U-shaped; margins entire, or sometimes slightly sinuate near apex; apex acuminate, acumen usually sharply differentiated, furrowed or strongly furrowed; costa double and short, very rarely single and ending well before mid leaf; alar region transversely triangular, reaching from margin 66-100% distance to costa; ratio of medial laminal cell length (µm) to leaf length (mm) 26-30.2. Sexual condition autoicous.
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