Plants medium-sized to large. Stems pinnate or irregularly branched. Stem leaves falcate or rarely almost straight, ovate, triangular, or broadly rounded-triangular, gradually narrowed to apex, concave, 1.5-6.4 × 0.5-1.3 mm; base erectopatent to patent, insertion slightly curved; margins entire or frequently partly finely denticulate; apex long-acuminate, acumen gradually differentiated, in straight-leafed plants ± plane; costa single, excurrent or longly so, rarely ending few cells before apex, strong; alar region transversely triangular or narrowly so, reaching from margin 66-80% distance to costa; ratio of medial laminal cell length (µm) to leaf length (mm) 16.5-23.9(-25.5). Sexual condition dioicous.