Plants medium-sized, sometimes small or large. Stems pinnate or irregularly pinnate. Stem leaves strongly falcate-secund, rarely weakly so, ovate, broadly ovate, or triangular-ovate, gradually narrowed to apex, ± strongly concave, 1.4-5.2 × 0.4-1.4 mm; base erect or erectopatent, insertion slightly curved; margins entire or weakly and obtusely denticulate; apex acuminate, acumen gradually differentiated, mostly furrowed; costa single, ending beyond mid leaf, mostly in acumen but not excurrent; alar region quadrate or transversely short-triangular, reaching from margin 40-60% distance to costa; ratio of medial laminal cell length (µm) to leaf length (mm) 23.3-36.5. Sexual condition dioicous.
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