Drymaria cordata Willd. ex Schult.

Whitesnow (en)

Species

Angiosperms > Caryophyllales > Caryophyllaceae > Drymaria

Characteristics

Glandular-puberulent to glabrate ramifying annuals, prostrate and spreading or erect, the internodes mostly longer than the nodes, glabrous to densely glandular, often rooting at the nodes. Leaves glabrous to scantily pubescent, orbiculate to reniform, apically rounded and occasionally mucronulate, basally rounded to cordate, 5-25 mm. long, 5-30 mm. broad, the petioles 2-15 mm. long; stipules mostly polylacerate, rather persistent, to 2 mm. long. Inflorescences of terminal or axillary few-many-flowered dichasial cymes (flowers rarely solitary in the axils) the bracteate pedicels locally girdled with a dense band of glandular pubescence, rarely subglabrous, 2145 mm. long, equaling to much exceeding the subtending bracts. Sepals 5, lanceolate to ovate, acute, glandular-puberulent to glabrous, obscurely to strongly 3-nerved, 2.5-4 mm. long, the borders translucent; petals 5, 2-3 mm. long, deeply bifid, the lobes linear, acute, rarely obtuse, 1-nerved, basally exauriculate but rarely subdentate; stamens 2-3 (-5), the flattened fila-ments 2-2.5 mm. long, the anthers suborbicular, 0.2-0.3 mm. long; styles 3, free nearly to their bases or united for half their length, 0.5-1 mm. long; ovules few to many, campylotropous on free central placentae. Capsule ovoid, 1.5-2.5 mm. long, the 3 valves entire; seeds 1-12, 1.0-1.5 mm. broad, cochleate, dark reddish brown, tuberculate in lines, all the tubercles low and domical, closely approximated or contiguous.
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Tender herb with stems straggling, procumbent or ascending, branching dichotomously, often rooting at the lower nodes, quadrangular, glabrous or papillose (especially above); leaves rather widely spaced. Leaves on glabrous petioles up to 1 cm long; lamina 1-3.5 x 0.6-3.0 cm, ovate to very broadly ovate, apex acute or subacute and shortly apiculate or occasionally rounded or obtuse, cordate, truncate or abruptly cuneate at the base, 3-7-nerved at the base, glabrous; stipules c. 1 mm long, interpetiolar, deeply fringed or composed of several subulate segments. Flowers in axillary or terminal, rather loose dichotomous cymes 2-25 cm long, peduncles slender and often elongate; bracts 1-4 mm long, narrowly lancolate, membranous; pedicels 1.5-5(12) mm long, filiform, glabrous or papillose. Sepals green with whitish margins, narrowly lanceolate, acute, keel viscid-papillose, with an additional nerve on each side of the midrib. Petals white, deeply bilobed, somewhat shorter than the sepals, sometimes absent. Stamens 5 or reduced to 3; filaments c. 2 mm long; anthers c. 0.5 x 0.4 mm Ovary ovoid-ellipsoid, usually 3-ovulate; style c. 0.25 mm long with 3 spreading stigmatic arms. Capsule c. 2 x 1.5 mm ellipsoid. Seeds 1-3, c. 1 x 0.75 mm, somewhat flattened-reniform; testa brown, minutely and bluntly tubercled.
A straggling herb with procumbent and more or less ascending branched stems, often rooting at the lower nodes, quadrangular, glabrous or papillose especially in the upper internodes; internodes slender, generally 2–6 cm. long.. Leaves opposite, blades ovate to very broadly ovate, acute or subacute and shortly apiculate or even rounded at apex, cordate, truncate, or shortly cuneate at base, 1 cm. long and 0.6 cm. broad up to 3.5 cm. long and 3.0 cm. broad, 3–7-palminerved, glabrous; petioles 2–7 mm. long; stipules inter-petiolar, about 1 mm. long, deeply fringed or composed of a number of subulate segments.. Cymes terminal or axillary, often loose on long slender peduncles and with slender branches; pedicels at anthesis mostly 3–5 mm. long.. Sepals narrowly lanceolate, acute, 3.5–5 mm. long, keeled along the papillose midrib, with one nerve on each side; the flowers (presumably the sepals) are said to be sticky.. Petals 3 mm. long, white.. Ovules usually about 3.. Seeds 1.5 mm. in greatest diameter, bluntly tubercled.. Fig. 4.
Annual herb to 45 cm long, erect to prostrate; stems branched, angled, often rooting at nodes, glabrous to densely glandular-puberulent. Petiole filiform, 2-15 mm long; stipules membranous, multilacerate, to 2 mm long; blade orbicular, ovate-orbicular or reniform, 5-25 x 5-30 mm, obtuse or acute and mucronulate at apex, round to cordate at base, glabrous or puberulent. Inflorescences terminal or axillary, few-flowered dichasial cymes, lax; pedicels 2-15 mm long, densely stipitate-glandular puberulent. Sepals 5, ovate, oblong-lanceolate or lanceolate, 2.5-3.5 mm long, glabrous except for stipitate-glandular puberulent midvein; petals 5, white, deeply clawed and bifid, shorter than calyx, 2-3 mm long, lobes linear; stamens 2-3(-5), filaments ca. 2 mm long; ovary ovoid, styles 3, 0.5-1 mm long. Capsule ovoid, slightly shorter than calyx, 1.5-2.5 mm long, shortly pedicellate; seeds 1-12, cochleate, 1.0-1.5 mm long, dark reddish-brown or black, densely tuberculate in lines.
Herbs, annual with fibrous roots, often adventitious from nodes, glabrous to densely pubescent, glandular or eglandular. Stems prostrate to ascending, to 30 (–60) cm long. Leaves petiolate, opposite or pseudoverticillate; petioles 2–15 mm long; lamina broadly ovate to reniform, obtuse, occasionally mucronate, 5–25 mm long, 5–30 mm wide; stipules lacerate to base, often in two layers, to 2 mm long. Inflorescence of branching, cymose panicles; bracts with broad, scarious margins, 1–5 mm long. Flowers 4–5 mm diam.; pedicels evenly glandular-puberulent, 1–8 mm long. Calyx ± glabrous. Sepals 2–4.5 mm long, thin to robust, prominently 1–3 (–7)-ribbed, ± carinate. Petals 5, 1.5–3 mm long, white, deeply bifid; lobes linear, acute to obtuse. Styles 3. Capsule 1.5–2.5 mm long, ovoid, 3-valved. Seeds 1–several, 0.8–2 mm wide, reddish brown, tuberculate.
Plants annual. Stems straggling, sometimes subscandent, usually rooting at lower nodes, 60--90 cm, mostly glabrous, minutely papillose toward inflorescence. Stipules membranous, splitting into few whitish setae. Petiole poorly defined, 3--7 mm; leaf blade ovate-cordate, (0.5--)1--3(--3.5) × 0.6--3 cm, prominently 3--5-veined from base. Bracts lanceolate, scarious. Pedicel 3--5 mm, slender, glandular hairy. Sepals lanceolate-ovate, 2--3.5(--5) mm, margin membranous, 3-veined, glandular hairy, apex subacute. Petals white, obovate-cuneate, ca. 2.5 mm, deeply 2-cleft; segments narrow, apex ± acute. Stamens 2--3(--5), shorter than sepals. Styles 3, connate at base. Capsule ovoid, (1.5--)2--3 mm in diam., 3-valved. Seed dark brown, suborbicular, ca. 1.5 mm, regularly densely tuberculate. Fl. Apr--Oct, fr. Jun--Dec.
Annual herb, 0.25-1.50 m high; procumbent or ascending, branching dichotomously, quadrangular, glabrous or papillose. Leaves petiolate, stipulate, glabrous, ovate to broadly ovate, rather widely spaced. Inflorescences axillary or terminal dichotomous cymes; flowers pedicellate. Sepals green with white margins, narrowly lanceolate, acute, keel viscid-papillose. Petals white, deeply bilobed, shorter than sepals, sometimes absent. Stamens 3-5. Ovary ovoid-ellipsoid. Flowering time Aug.-Apr. Fruit a capsule, ellipsoid. Seeds flattened-reniform, testa brown, minutely or bluntly tubercled.
An annual herb. It climbs or creeps along the ground. It has branches that divide in two. It often forms a dense ground cover. The stems form roots at the nodes. The leaves are small, opposite and heart shaped. The leaves are 0.8-2 cm long by 0.5-2 cm wide. The flowers are small and white. The are in twice branched heads at the ends of the stems. The fruit is a capsule. It splits to release small seeds.
Leaves on glabrous petioles up to 1 cm. long; lamina 1–3·5 × 0·6–3·0 cm., ovate to very broadly ovate, apex acute or subacute and shortly apiculate or occasionally rounded or obtuse, cordate, truncate or abruptly cuneate at the base, 3–7-nerved at the base, glabrous; stipules c. 1 mm. long, interpetiolar, deeply fringed or composed of several subulate segments.
Flowers in axillary or terminal, rather loose dichotomous cymes 2–25 cm. long, peduncles slender and often elongate; bracts 1–4 mm. long, narrowly lancolate, membranous; pedicels 1·5–5 (12) mm. long, filiform, glabrous or papillose.
Tender herb with stems straggling, procumbent or ascending, branching dichotomously, often rooting at the lower nodes, quadrangular, glabrous or papillose (especially above); leaves rather widely spaced.
Sepals green with whitish margins, narrowly lanceolate, acute, keel viscid-papillose, with an additional nerve on each side of the midrib.
Seeds 1–3, c. 1 × 0·75 mm., somewhat flattened-reniform; testa brown, minutely and bluntly tubercled.
Ovary ovoid-ellipsoid, usually 3-ovulate; style c. 0·25 mm. long with 3 spreading stigmatic arms.
Petals white, deeply bilobed, somewhat shorter than the sepals, sometimes absent.
Stamens 5 or reduced to 3; filaments c. 2 mm. long; anthers c. 0·5 × 0·4 mm.
Capsule c. 2 × 1·5 mm. ellipsoid.
Life form annual
Growth form herb
Growth support -
Foliage retention
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.5 - 0.6
Root system adventitious-root fibrous-root
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

A tropical plant. It grows in damp shaded sites, often near streams or under shrubs, disturbed areas between 200–1900(–2400) m altitude in China. It grows in wetlands. It grows in Nepal between 2200-4300 m altitude. In Tanzania it grows between 900-2,700 m above sea level. It grows in open areas. In XTBG Yunnan. In Sichuan.
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Grassland, forest margins, roadsides and cultivated areas, often under shade, at mid to higher elevations. The plant appears spontaneously as a weed of cultivation.
Lowland in C. Japan. The plant grows in tropical and sub-tropical India, but extends into the Himalayas up to elevations of 2100 metres.
Weed of disturbed agricultural and urban areas.
Light 7-9
Soil humidity 4-6
Soil texture 1-6
Soil acidity 3-7
Soil nutriment -
Hardiness (USDA) 9-12

Usage

The tender shoots and leaves are cooked as a vegetable. They are also eaten in salads. The shoots can be stored for 4-5 days.
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Use: The plant is used in a medicinal bath for children in French Guiana.
Uses animal food environmental use erosion control fodder food gene source material medicinal social use
Edible leaves roots shoots
Therapeutic use Anti-bacterial agents (aerial part), Abortifacient agents (leaf), Burns (leaf), Epistaxis (leaf), Headache (leaf), Sinusitis (leaf), Skin diseases (leaf), Tonsillitis (leaf), Urination disorders (leaf), Wounds and injuries (leaf), Apertif (unspecified), Boil (unspecified), Depurative (unspecified), Fever (unspecified), Laxative (unspecified), Medicine (unspecified), Poultice (unspecified), Stimulant (unspecified), Tumor (unspecified), Emollient (unspecified), Antifungal agents (unspecified), Anti-inflammatory agents (unspecified), Antitussive agents (unspecified), Burns (unspecified), Common cold (unspecified), Cough (unspecified), Diarrhea (unspecified), Dysentery (unspecified), Dysmenorrhea (unspecified), Fractures, bone (unspecified), Furunculosis (unspecified), Headache (unspecified), Influenza, human (unspecified), Laxatives (unspecified), Mouth diseases (unspecified), Nasal disease (unspecified), Peptic ulcer (unspecified), Sinusitis (unspecified), Skin diseases (unspecified), Snake bites (unspecified), Urination disorders (unspecified), Vomiting (unspecified), Wounds and injuries (unspecified), Fever (whole plant), Laxatives (whole plant), Cooling effect on body (whole plant)
Human toxicity -
Animal toxicity -

Cultivation

Plants are grown from seed.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Leaf

Drymaria cordata leaf picture by Donna Shaynak (cc-by-sa)
Drymaria cordata leaf picture by Paulo Cardoso (cc-by-sa)
Drymaria cordata leaf picture by Teixeira Wenceslau (cc-by-sa)

Flower

Drymaria cordata flower picture by Teixeira Wenceslau (cc-by-sa)
Drymaria cordata flower picture by nicoplant (cc-by-sa)

Fruit

Drymaria cordata fruit picture by Frederick Henderson (cc-by-sa)

Distribution

Drymaria cordata world distribution map, present in Argentina, Antigua and Barbuda, Australia, Bonaire, Sint Eustatius and Saba, Bahamas, Belize, Bolivia (Plurinational State of), Brazil, Barbados, Chile, China, Colombia, Costa Rica, Cuba, Dominica, Ecuador, Fiji, Guadeloupe, Grenada, Guatemala, French Guiana, Guyana, Honduras, Jamaica, Japan, Saint Lucia, Mexico, Montserrat, Martinique, Nicaragua, Panama, Peru, Puerto Rico, Paraguay, El Salvador, Suriname, Trinidad and Tobago, Uruguay, United States of America, Venezuela (Bolivarian Republic of), and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:154112-1
WFO ID wfo-0000657240
COL ID 6DLHG
BDTFX ID -
INPN ID 447070
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Alsine rotundifolia Bufonia rotundifolia Drymaria diandra Holosteum cordatum Alsine media Drymaria adenophora Drymaria procumbens Drymaria sessilifolia Holosteum diandrum Stellaria adenophora Drymaria cordata var. diandra Drymaria cordata var. pacifica Drymaria cordata var. puberula Drymaria cordata subsp. cordata Drymaria cordata var. cordata Drymaria cordata

Lower taxons

Drymaria cordata subsp. diandra