Dryopteris fragrans (L.) Schott

Fragrant woodfern (en), Fougère-mâle (fr)

Species

Pteridophytes > Polypodiales > Dryopteridaceae > Dryopteridoideae > Dryopteris

Characteristics

Plants 20-30 cm tall. Rhizome erect or obliquely ascending, apex and stipe base scaly; scales ferruginous, ovate or ovate-lanceolate, membranous, sparsely dentate, shortly acuminate. Fronds caespitose; stipe stramineous, 1-2(-12) cm; scales ferruginous, oblong-lanceolate, with capitate golden glands, dentate; lamina oblong-lanceolate, 10-25 × 2-4 cm, widest at middle, bipinnate, gradually narrowed toward base, base less than 1 cm wide, apex shortly acuminate; pinnae ca. 20 pairs, oblique, lanceolate, apex obtuse or abruptly acuminate, middle pinnae 1.5-2 × 6-8 cm, widest at base, lower several pairs narrowed to auricles; pinnules oblong, dentate or lobed. Lamina herbaceous, glabrous on both surfaces; rachis and costa clothed with lanceolate, glossy, brown scales and glands; veins pinnate, obscure on both surfaces. Sori abaxial on veinlets; indusia orbicular or orbicular-reniform, membranous, sparsely dentate, close to each other at maturity and always exserted from frond margin, glandular; spores elliptic, tuberculate on epispore. 2n = 82.
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Leaves monomorphic, green through winter, 6--40 × 1--6 cm, old leaves persistent as gray or brown conspicuous clump at plant base (only in this species). Petiole 1/3 length of leaf, scaly throughout; scales dense, brown to red-brown. Blades green, linear-lanceolate, pinnate-pinnatifid to 2-pinnate, papery, densely scaly (only in this species) abaxially, glandular, aromatic when handled (only in this species). Pinnae ± in plane of blade, linear-oblong; basal pinnae linear-oblong, much reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal; pinnule margins crenately toothed. Sori midway between midvein and margin of segments. Indusia glandular. 2 n = 82.
Plants Woodsia-like in aspect; rhizomes ± erect; lvs evergreen (and long-persistent after death), firm, fragrant, glandular and scaly; petiole short, to 7 cm; blade lance-linear, pinnate-pinnatifid, to 30 × 5.5 cm, acuminate, gradually narrowed below, the lowest pinnae very small; pinnae numerous and crowded, mostly 15–25 pairs, linear-oblong, to 27 × 8 mm, with 6–10 pairs of oblong, obtuse, crenate segments to 5 mm, not spinulose; veins obscure, 2–3 pairs per segment; sori large, often only 1–4 per segment; indusia glandular-margined; 2n=82. Cliff-crevices, often on limestone; interruptedly circumboreal, s. to N. Y., Ont., n. Mich., Wis., and Minn.
A fern. It grows 20-30 cm tall. The blade is oblong to sword shaped and 10-25 cm long by 2-4 cm wide.
Life form perennial
Growth form herb
Growth support -
Foliage retention evergreen
Sexuality -
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.2
Root system rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Rocks and screes in Arctic Finland. Shaded cliffs and talus, often of limestone at elevations of 50-1800 metres in northern N. America.
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It is a temperate plant. In north China it grows between 700-2,400 m above sea level.
Light 4-6
Soil humidity 5-7
Soil texture 3-5
Soil acidity 3-6
Soil nutriment -
Hardiness (USDA) 3-6

Usage

Uses environmental use fiber medicinal tea
Edible leaves
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Dryopteris fragrans unspecified picture

Distribution

Dryopteris fragrans world distribution map, present in Canada, China, Finland, Japan, Kazakhstan, Liberia, Mongolia, Korea (Democratic People's Republic of), Russian Federation, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:17095220-1
WFO ID wfo-0001107303
COL ID 37WFT
BDTFX ID 83211
INPN ID 114965
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Filix fragrans Filix-mas fragrans Dryopteris fragrans Aspidium fragrans Polypodium fragrans Lastrea fragrans Nephrodium fragrans Woodsia xanthosporangia Dryopteris aquilonaris Dryopteris fragrans var. remotiuscula Dryopteris fragrans var. fragrans Nephrodium fragrans f. aquilonare Dryopteris fragrans f. aquilonaris Filix aquilonaris Nephrodium fragrans var. remotiusculum Thelypteris fragrans var. hookeriana