Dryopteris squamiseta (Hook.) Kuntze

Plants terrestrial. Rhizome erect to suberect, usually unbranched, to 60 mm long and 8 mm in diameter, closely set with roots, stipe bases, and scales, the scales chartaceous, castaneous, adnate, narrowly triangular to subulate, to 11 x 2 mm, the margins regularly set with short multicellular, uniseriate hairs and capitate glands, the glands also occur on the scale lamina, the scale apex terminates in a short uniseriate series of cells. Fronds up to 8 per plant, suberect to arching, to 925 mm long; stipes firm, proximally castaneous, brown to stramineous higher up, adaxially sulcate, to 460 mm long and 5 mm in diameter, closely set with patent scales, hairs, and glands, the scales chartaceous to thinly crustaceous, castaneous, adnate, narrowly triangular to subulate, to 11 x 2 mm, entire, or those near the base often with short multicellular, uniseriate hairs and capitate glands, the glands often also occur on the scale lamina, the scale apex terminates in a short uniseriate series of cells, the hairs are of 2 kinds: 1) inconspicuous, pale, multicellular uniseriate appressed hairs and, 2) castaneous, multicellular uniseriate patent hairs, the cells collapsing in the ctenitoid fashion upon drying, the glands clavate; laminae proximally anadromous, catadromous towards the apex, to 3-pinnate, ovate, to 460 mm long, to 390 mm wide, with up to 10 petiolated pinna pairs; rachises stramineous, adaxially sulcate, the sulcus open to that of the pinna-rachis, closely set with scales similar to, but smaller than those on the stipe; pinnae petiolate, the petiole up to 13 mm long, opposite to alternate, overlapping, the basal pinna pair strongly basiscopically developed, inaequilaterally narrowly ovate, lanceolate to oblong-acute towards the apex, to 220 mm long, to 125 mm wide, 1-pinnate-pinnatifid to 2-pinnate, with up to 5 free pinnule pairs; pinna-rachises stramineous, adaxially shallowly sulcate, moderately scaled, the scales patent, chartaceous, castaneous, adnate, subulate, to 3 mm long; pinnules petiolate, the petiole to 2 mm long, catadromous, alternate, spaced or overlapping, oblong-acute to oblong-attenuate, acroscopic pinnule on basal pinnae to 44 mm long, to 17 mm wide, basiscopic pinnule on basal pinnae to 82 mm long, to 24 mm wide, pinnatifid to 1-pinnate; segments herbaceous, trapeziform to inaequilaterally oblong-obtuse, often acroscopically auricled, basiscopically decurrent along the costa towards the pinnule apex, lobed to crenulate, up to 14 x 6 mm, adaxially with simple, proximally multistratose subulate hairs along the veins, to 1.2 mm long, abaxially with proximally multistratose subulate hairs and clavate, pluricellular, uniseriate hairs, up to 0.85 mm long, and capitate glands to 30 µm long along the veins. Venation anadromous, evident, pinnately branched, free vein branches end near the margin, fertile vein branches undifferentiated or variously shortened beyond the sorus. Stomata of the anomo-and polocytic types, guard cells (32-)48(-64) µm long. Sori circular, usually restricted to the pinnule and segment apices, medially to inframedially on unmodified vein branches, to 1.5 mm in diameter; indusia firmly herbaceous, brown, persistent, reniform to subcircular, entire, to 1.2 mm in diameter, adaxially with numerous capitate glands and rarely also with a few short hairs near the point of attachment; sporangia long-stalked, the stalk simple or with a long uniseriate trichome, 3-seriate below the capsule, the capsule globose, with 13(-15) indurated annulus cells, epistomium 4(-5)-celled, hypostomium 4(-5)-celled. Spores 64 per sporangium, brown, ellipsoidal, monolete, with prominent folds, tuberculate, (32-)35(-36) x (18-)20(-22) µm.