Heads radiate, the rays pistillate, usually white, very short; invol bracts 1–2-seriate, herbaceous or herbaceous-tipped, subequal, or the inner narrower and shorter; receptacle flat or slightly convex, its bracts slender and fragile, or the inner ones wanting; disk-fls perfect and fertile, 4(5)-toothed; style-branches flattened, with short, obtuse, hairy appendage; achenes thick, commonly transversely rugose, 3–4-angled, those of the disk somewhat compressed at right angles to the invol bracts; pappus none or an obscure crown or 2 short awns; branching herbs with opposite simple lvs. 4, mostly tropical.
Annual herbs. Leaves opposite. Capitula axillary or terminal, solitary or in pairs or in panicles, heterogamous, radiate; involucre 2-seriate; receptacle paleate. Ray florets female or sterile, with a small ray; disc florets hermaphrodite, tubular to campanulate, 4–5-lobed; anthers with rounded terminal appendage; style-arms obtuse. Achenes ± compressed, rugose to tuberculate; pappus absent or of 2 short awns.