Herbs perennial, monoecious or dioecious, 25-90 cm tall. Stems ascending or erect, simple or rarely branched, lower stems ferruginous-furfuraceous, glabrous. Leaves alternate; nanophylls absent; stipules linear or linear-lanceolate, 5-10(-20) × 2-3 mm, glabrous, with cystoliths; petiole 0-3 mm; leaf blade obliquely elliptic or oblong, 2-13.5(-23) × 1-5(-8) cm, herbaceous, major basal lateral veins asymmetric, 1 basal, the other arising above base, both surfaces glabrous or adaxial surface sparsely strigillose; cystoliths conspicuous, dense; base with broader half broadly cuneate or nearly auriculate, margin dentate, apex cuspidate. Male inflorescences solitary, simple, 6-12 mm in diam.; peduncle 1.5-4 mm; receptacle oblong, 6-11 × 5-7 mm; bracts connate, 0.5-1.6 mm; bracteoles oblong or cymbiform-oblong. Female inflorescences solitary, 3-15 mm in diam.; peduncle 0.5-1 mm; receptacle 2-13 mm in diam.; bracts broadly ovate or triangular; bracteoles narrowly linear. Male flowers 4-merous. Achenes narrowly ellipsoid, 8-ribbed. Fl. May-Oct.
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