Herbs perennial or subshrubs, monoecious or dioecious, 60-200 cm tall. Stems erect, branched, glabrous or strigillose. Leaves alternate, glabrous; nanophylls absent; stipules narrowly lanceolate, 6-10 × 1.5-2 mm, with cystoliths; petiole 1.5-6(-10) mm; leaf blade obliquely elliptic to obliquely oblanceolate, 5-19 × 2-6 cm, papery or herbaceous, major basal lateral veins asymmetric, 1 basal, the other arising above base, cystoliths conspicuous, dense; base obliquely cuneate, margin entire or ca. 2-crenate, apex acuminate or caudate. Male inflorescences solitary or in pairs, simple, 10-18 × 6-12 mm, sessile; receptacle 4-6 mm in diam.; bracts connate, ca. 1 mm; bracteoles cymbiform. Female inflorescences solitary or in pairs, 5-8 mm in diam.; peduncle 0-1 mm; receptacle very small; bracts triangular; bracteoles narrowly linear. Male flowers 4-merous. Achenes ellipsoid, 8-ribbed. Fl. Mar-May.