Herbs perennial, monoecious or dioecious, 20-150 cm tall. Stems ascending or erect, simple or branched, lower stems ferruginous-furfuraceous, glabrous. Leaves alternate; nanophylls absent; stipules lanceolate, 13-17 × 4-6 mm, glabrous, with cystoliths; petiole 2-6(-15) mm; leaf blade obliquely elliptic or oblong, 9-25(-28) × 3.5-9 cm, papery, major basal lateral veins both arising at base of leaf blade, both surfaces glabrous or adaxial surface sparsely puberulent; cystoliths conspicuous, dense; base obliquely cuneate, margin dentate, apex acuminate or caudate-acuminate. Male inflorescences solitary or rarely in pairs, simple, 7-18 × 5-9 mm in diam.; peduncle 1-3 mm; receptacle elliptic, 6-12 × 5-8; bracts connate, 0.5-1 mm; bracteoles linear. Female inflorescences solitary, 9-13 × 6-9 mm in diam.; peduncle 0.5-1 mm; receptacle elliptic, 7-11 × 4-7 mm; bracts broadly ovate; bracteoles spatulate. Male flowers 4-merous. Achenes narrowly ellipsoid, 9-ribbed. Fl. Apr-Jun.