Herbs perennial, monoecious or dioecious, 25-50 cm tall. Stems ascending or erect, simple or branched, crispately pubescent. Leaves alternate; nanophylls cordate or ovate, 3-10 mm; stipules lanceolate-linear, 5-7 × 0.5-0.8 mm, glabrous, without cystoliths; petiole 1-5 mm; leaf blade obliquely elliptic or obliquely obovate, 4.5-13.5 × 2-5.5 cm, papery, major basal lateral veins both arising at base of leaf blade or 1 arising above base, not extending beyond middle of leaf blade, abaxial surface puberulent along nerves, adaxial surface glabrous, cystoliths conspicuous or obscure, dense; base with broader half broadly cuneate or nearly auriculate, margin crenate, apex cuspidate or caudate. Male inflorescences solitary, simple, 11-26 × 10-15 mm; peduncle 20-55 mm; receptacle elliptic, 10-25 × 10-14 mm; bracts connate, 5-12 mm wide; bracteoles spatulate-linear. Female inflorescences solitary, 4-6 mm in diam.; peduncle ca. 1 mm; receptacle nearly orbicular, 3-5 mm in diam.; bracts triangular; bracteoles lanceolate-linear. Male flowers 4-merous. Fl. Jun-Jul.