Herbs perennial, monoecious, 20-40 cm tall, glabrous. Stems ascending or erect, branched. Leaves alternate; nanophylls absent; stipules narrowly triangular, 0.7-1.5 × 0.2-0.4 mm, without cystoliths; petiole 1-3 mm; leaf blade obliquely ovate or elliptic-ovate, 4-11 × 1.5-4 cm, membranous or herbaceous, major basal lateral veins both arising at base of leaf blade, cystoliths conspicuous, dense; base obliquely cuneate, margin denticulate, apex acuminate or cuspidate. Male inflorescences solitary, simple, 4-7 mm in diam.; peduncle 12-40 mm; receptacle small; bracts connate, broadly ovate. Female inflorescences solitary, 30-flowered or more, 3-6 mm in diam., sessile; receptacle 3-5 mm in diam.; bracts narrowly ovate; bracteoles spatulate-linear. Male flowers 4-merous. Achenes ellipsoid, smooth. Fl. Sep-Oct.