Herbs perennial, monoecious or dioecious, 15-50 cm tall. Stems ascending or erect, simple or 1-branched, glabrous or pilose. Leaves alternate; nanophylls absent; stipules subulate, 0.6-1 × 0.1 mm, glabrous, without cystoliths; petiole 0.5-2 mm; leaf blade obliquely elliptic-ovate or elliptic, 2-5 × 1.2-2 cm, herbaceous, major basal lateral veins asymmetric, 1 basal, the other arising above base, abaxial surface glabrous, adaxial surface sparsely strigose, cystoliths obscure, sparse; base with broader half cordate or nearly auriculate, margin denticulate, apex acute or nearly obtuse. Male inflorescences solitary, simple, 5-10 mm in diam., sessile; receptacle obscure; bracts connate, to 5 mm. Female inflorescences solitary, 30-flowered or more, 2-5 mm in diam.; peduncle 2-12(-35) mm; receptacle small; bracts connate, narrowly ovate or narrowly lanceolate; bracteoles lanceolate-linear. Male flowers 4-merous. Achenes narrowly ovoid, ribbed, tuberculate. Fl. May-Aug.