Eleocharis retroflexa Urb.

Coastal plain spikerush (en)

Species

Angiosperms > Poales > Cyperaceae > Eleocharis

Characteristics

Plants annual, tufted, mat-forming, often stoloniferous, sometimes entirely vegetative; rhizomes absent. Culms erect, ascending or arching, pentagonal, sulcate, 1.5–10 cm × 0.2–0.3 mm [larger], soft. Leaves: distal leaf sheaths persistent or disintegrating, pale brown to green, red-spotted [mostly red-brown], membranous; apex acuminate. Spikelets: basal spikelets usually present, bisexual; often proliferous, ellipsoid or obovoid, laterally compressed, 1.7–3.9 × 1.2–2 mm, apex acute; proximal scale empty or with a flower, deciduous, amplexicaulous, similar to floral scales (sometimes 2.4–2.9 mm); subproximal scale with a flower; floral scales clearly distichous, 2–6 [or more], 4–6 per mm of rachilla, pale brown [marked red-brown], ovate or elliptic, 1.8–2.5 × 0.8–1.4 mm, membranous, apex rounded to obtuse, midribs green, keeled. Flowers: perianth bristles 6, colorless or pale brown, shorter than achenes; spinules not evident at 45X; stamens 3; anthers (0.55–)0.7 mm; styles 3-fid. Achenes stramineous (to cream), obovoid, trigonous or subterete, not compressed, angles prominent, 0.8 × 0.5–0.55 mm, apex not constricted proximal to tubercle, coarsely cancellate or honeycomb-reticulate at 10–15X. Tubercles red-brown, pyramidal, trigonous, proximally clearly to obscurely 3-lobed, lobes decurrent on achene angles, 0.3–0.35 × 0.3–0.4 mm.
More
Annual. Stems very slender, tufted, often curved, angular, 4-5-ribbed, puncticulate, light green, 5-20 cm by 1/5-⅓ mm. Sheaths reddish at the base, whitish-scarious at the apex. Spikelet ovoid, obtuse, more or less compressed, loosely few-(3-10)-flowered, 2-5 by 1½-2½ mm. Glumes membranous, spreading in fruit, ovate, obtuse, keeled, with distinct greenish midnerve, purplish sides, and scarious margins, 2¼-3¼ by. 1½-2 mm, the lower ones almost distichous. Bristles 6, rather delicate, unequal, the longest usually longer than the nut, retrorsely scabrous, whitish to yellowish, occasionally reduced to obsolete. Stamens 3; anthers linear, minutely apiculate, ½-¾ Style 3-fid. Nut equilaterally triquetrous, urceolate, truncate or slightly convex at the apex, conspicuously cancellate, costulate on the angles, dull stramineous to fumose, 9/10 by 7/10-9/10 mm; epidermal cells conspicuous, deeply pitted, hexagonal or roundish, in 6-10 vertical series on each face. Style-base pyramidal, in the Malesian specimens usually much depressed and blunt at the apex, c. ⅓ as high as and equal in width to the nut, 3-lobed, the lobes decurrent on the angles of the nut.
Annuals. Roots fibrous. Rhizomes and stolons absent. Culms often recurved, tufted, 2-16 cm tall or rarely more, filiform, soft, 5-or nearly 4-angled. Leaf sheaths 1 or 2, pale red, tubular, 2-12 mm; basalmost sheath bladeless; cauline sheaths usually with a glumelike blade at apex. Spikelet purplish red, ovoid, ca. 4 × 2 mm, slightly compressed, few flowered, sometimes proliferous. Basalmost glume empty, subelliptic, amplexicaul for whole spikelet base; fertile glumes green, broadly ovate, 2.8-3 × 1.8-2 mm, herbaceous, with a median keel, edge membranous, margin hyaline, apex acute to obtuse. Perianth bristles 6, nearly as long as nutlet, one on outer series shorter than nutlet, retrorsely spinulose. Stigmas 3. Nutlet yellowish, broadly obovoid, ca. 1.5 × 1 mm, 3-sided, angles prominently costate, reticulate to deeply pitted-reticulate with isodiametric epidermal cells, apex broadly truncate and not constricted; persistent style base pyramidal-acuminate, base as wide as to slightly narrower than nutlet. Fl. and fr. Jul-Nov.
Rhizomes filiform; culms capillary, 2-20 cm. long; spikelets 3-4 mm. long, containing 1-4 achenes, green, sometimes spotted with purple, the scales acutish; bristles equaling the achene, which is obovoid, trigonous, white, coarsely cancellate, the tubercle pyramidal.
Life form annual
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.02 - 0.13
Root system fibrous-root rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway -

Environment

In wet muddy places: along streams, along humid road-sides, in shallow pools, in rice-fields, 0-1600 m.
Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 9-12

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Distribution

Eleocharis retroflexa world distribution map, present in Angola, Åland Islands, Andorra, Australia, Bangladesh, Belize, Bolivia (Plurinational State of), Brazil, Botswana, Central African Republic, China, Congo, Costa Rica, Cuba, Dominica, Ecuador, Gabon, Guinea, Guatemala, Guyana, Honduras, Indonesia, Jamaica, Liberia, Moldova (Republic of), Mali, Myanmar, Mauritius, Nicaragua, Nepal, Panama, Peru, Philippines, Puerto Rico, Sierra Leone, Suriname, Thailand, Tanzania, United Republic of, United States of America, Venezuela (Bolivarian Republic of), Viet Nam, Wallis and Futuna, South Africa, and Zambia

Conservation status

Eleocharis retroflexa threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:89818-2
WFO ID wfo-0000405873
COL ID 397LS
BDTFX ID -
INPN ID 629611
Wikipedia (EN)
Wikipedia (FR)

Synonyms

Baeothryon retroflexum Eleocharis retroflexa Scirpus radiciflorus Scirpus retroflexus Chaetocyperus niveus Chaetocyperus polymorphus var. depauperatus

Lower taxons

Eleocharis retroflexa subsp. chaetaria Eleocharis retroflexa subsp. retroflexa Eleocharis retroflexa subsp. subtilissima