Trees 5-50 m. Twigs (pale) grey brown to dark brown, 2-6(-10) mm diameter, with grey-brown to rusty hairs 0.1-0.5 mm, early to late glabrescent, bark (coarsely) striate or longitudinally cracking, sometimes flaking, lenticels absent or inconspicuous. Leaves membranous or chartaceous, elliptic or obovate to oblong-oblanceolate, (12-) 15-35 (-40) by (4-)5-12 cm, base (narrowly) rounded to attenuate, apex acute-acuminate; upper surface olivaceous to brown, glabrous, lower surface glabrous or (early) glabrescent; midrib above slender, flat or slightly raised; nerves 11-24 pairs, lines of interarching generally indistinct; venation lax or fine, sometimes slightly trabeculate, on both surfaces distinct or not; petiole 10-25 by 1.5-3.5 mm, glabrous to late glabrescent; leaf bud 8-25 by 1.5-3.5 mm, with dense grey brown to rusty hairs 0.1-0.5 mm. Inflorescences (see note 1) slender to stout, much or little branched, glabrescent or with persistent greyish or rusty hairs 0.1-0.5 mm long, in male 6-30 by 2-25 cm, moderately to many-flowered, peduncle 0-5 cm long, bracts ± elliptic, 1.5-3 mm, ± thinly pubescent, caducous. Flowers 4-10 together, buds usually in different stages of development, in ± umbelliform clusters (3-)5-30 mm spaced along the main branches of the inflorescence; perianth outside early glabrescent or thinly with hairs 0.1-0.5 mm long, lobes inside towards the apex with few to many (pale) brownish red hairs 0.2-0.4 mm long, usually in rows in spaces between the anthers, sometimes only near the lobe sutures. Male flowers: pedicel 1.5-7 mm long; buds broadly ellipsoid or broadly ovoid, 1.5-2.3 by 1.3-2 mm, apex rounded to subacute, base rounded to attenuate, in cross section sub-circular or 3-or 4-angular, cleft 2/3-4/5, lobes 3 or 4 (or 5), at sutures 0.2-0.3(-0.5) mm thick; synandrium (depressed) globose to ellipsoid, 0.2-0.5 by 0.3-0.5 mm, anthers 3-6, androphore slender, longer to shorter than the androecium, 0.3-1 mm long. Female flowers: pedicel 2-6 mm; buds ovoid-ellipsoid, 2.2-2.6 by 1.7-2 mm, cleft 2/3-5/6, lobes 0.2-0.3 mm thick; ovary (narrowly) ovoid, 1.3-1.8 by 0.8-1.2 mm, glabrous, stigma 0.2-0.3 mm high, narrowly to broadly 2-lipped, in the latter case the lips minutely 2-5-lobulate. Fruits 3-12 in a pendent loose panicle (10-) 15-30 cm long, ovoid, (narrowly) ellipsoid, or obovoid, 1.5-4.5(-5.5) cm long, glabrous; pericarp 1-3 mm thick, blackish, finely granulate, not or sparingly warted; the aril almost entire to laciniate to about halfway; seeds usually with pointed apex, testa variegated (not always in New Guinea).
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