Herbs perennial, erect. Basal leaves largest, margin coarsely serrate to incised-pinnatipartite. Verticillasters many flowered, widely spaced or in long robust spikes, densely lanate-villous or glabrous. Flowers sessile. Calyx tubular-campanulate to broadly funnelform, 5-toothed, sometimes limb dilated, 5-apiculate; teeth short, broadly truncate, rounded, ovate or triangular, apex spiny; sinuses between teeth with ovate-triangular appendages. Corolla 2-lipped; tube mostly included, narrow; upper lip narrow, galeate or falcate, concave, narrowed at base, bearded or villous inside and on margin; lower lip spreading, 3-lobed, middle lobe largest. Stamens 4, anterior 2 longer; some filaments with comblike-fimbriate appendages at base; anther cells 2, divaricate. Style unequally 2-cleft, anterior lobe larger. Nutlets obovoid, triquetrous, subtruncate, densely bearded at apex.
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