Trees or shrubs. Leaves alternate; leaf blade oblong-lanceolate, densely pinnately veined, apex acute. Inflorescence axillary, long pedunculate, capitate, with involucre; bracts 2-4, caducous. Flowers bisexual, sessile, 4-merous. Calyx tube white, funnel-shaped, exterior densely pilose; lobes ± unequal; imbricate. Petaloid appendages absent. Stamens twice as many as calyx lobes, in two series; filaments very short; anthers linear, introrse, basifixed; upper ones half exserted from throat. Disk cup-shaped, membranous, unequal, lobed or serrate, sometimes developed on one side. Ovary ellipsoid, 1-loculed; style short; stigma capitate, included. Fruit a berrylike drupe. Seed testa crustaceous.