Perennials, 40–100+ cm. Stems (from short rhizomes) single, densely branched distally, puber-ulent throughout. Leaves usually opposite (distal sometimes alternate, lateral buds dormant or producing 1 pair of leaves); simple, sessile or subsessile; blades ± 3-nerved distal to bases, elliptic to lance-elliptic, (30–)50–70 × 5–25 mm (lengths mostly 2–5 times widths), bases narrowly cuneate, margins usually serrate, apices acute, faces puberulent or villous, densely gland-dotted. Heads in corymbiform arrays. Phyllaries 7–10 in 2–3 series, elliptic, 1–3 × 0.5–1 mm, apices rounded to acute (not mucronate), abaxial faces puberulent, gland-dotted. Florets 5; corollas 2.5–3 mm. Cypselae 1.5–2 mm; pappi of 30–40 bristles 2.5–3 mm. 2n = 20.