Plants to 6 × 1 mm. Stem branched and unbranched; axil-lary hyaline nodules absent; central strand weak or absent. Leaves as many as 18 pairs, lingulate to lanceolate, rounded to obtuse to broadly acute, to 1 × 0.25 mm; dorsal lamina narrowed or rounded proximally, ending at insertion, not decurrent; vaginant laminae ± 1/2-2/3 leaf length, ± unequal, minor lamina ending near margin; margin serrulate, limbate, ± entire in proximal 1/3-1/2 of perichaetial and subtending 1-2 pairs of leaves, limbidium sometimes indistinct, limbidial cells 1-stratose; costa ending 3-25 cells before apex, often spurred distally, bryoides-type; laminal cells 1-stratose, pluripapillose, obscure, firm-walled, irregularly quadrate to hexagonal, 4-8 µm, in transverse section usually twice as deep as wide. Sexual condition rhizautoicous, cladautoicous, rarely gonioautoicous. Sporophytes 1-2 per peri-chaetium. Seta to 2 mm. Capsule theca exserted, erect, radially symmetric, to 0.6 mm; peristome scariosus-type; operculum 0.3 mm. Calyptra cucullate, ± prorate, 0.3 µm. Spores 9-14µm.