Annual or perennial herbs or shrubs. Stems erect, strongly branched, glandular-pubescent and/or with appressed non-glandular hairs. Leaves simple, tripartite or biternately dissected, opposite at first, becoming alternate, ovate to narrowly lanceolate, entire or dentate. Inflorescence cymose, subcorymbose. Capitula discoid, homogamous, pedunculate; involucral bracts uniseriate, acute or obtuse, usually scarious on margins. Florets all bisexual; corolla white to purplish, noticeably zygomorphic; style branches flattened, recurved, terminated by a penicillate cuspidate appendage. Achenes shortly pubescent, sometimes glabrescent, obpyramidal, 4-sided, with faces smooth or ribbed. Pappus persistent, of 8–10 ovate to truncate scales, with midvein rarely extending to apex or beyond.