Shrubs (0.5–)1–2.5 m, branchlets puberulent, glabrescent. Leaves: blade gray-green, flat to concave-convex, elliptic to oblong-elliptic, ovate-elliptic, or broadly lanceolate, 1.6–4(–5.5) × 0.7–2.5 cm, length 2 times width, margins undulate, ± muricate-roughened <and with callose rim>, especially distally, apex obtuse to subacute and mucronulate, both surfaces usually persistently densely tomentulose, <hairs coiling to recurved>, or adaxial sometimes barely glossy, ± glabrescent. Aments: staminate 2–3 cm; pistillate loose, internodes 4+ mm, sometimes branched, erect, 1.5–3 cm; pistillate bracts distinct or connate basally, each usually subtending 1 flower, elliptic to ovate, at least proximal similar in size and shape to distal leaves, closely puberulent with curly or crisped hairs. Berries 4–8 mm diam., glabrous, not glaucous.