Shrubs or trees 1–3.5(–5) m, branchlets puberulent, glabrescent. Leaves: blade green, flat, oblong-elliptic to broadly elliptic or obovate, 4.5–8 × 2.5–5 cm, length 2 times width, margins flat, smooth, <without callose rim>, apex rounded and mucronulate, abaxial surface persistently sparsely to densely puberulent-tomentulose, <hairs coiling to recurved>, adaxial surface glossy, glabrous or glabrate. Aments: staminate 2–3 cm; pistillate loose, internodes 4+ mm, sometimes branched, erect, 2–8 cm; pistillate bracts distinct or connate basally, each usually subtending 1 flower, elliptic to ovate, at least proximal similar in size and shape to distal leaves, minutely puberulent, <hairs tightly coiling to strongly recurved>. Berries 5–10 mm diam., glabrous, usually glaucous. 2n = 22.