Trees. Leaves alternate; seedling leaves simple, dentate; intermediate leaves pinnatisect to imparipinnate to bipinnate; adult leaves simple to bipinnate; margins dentate. Conflorescence lateral or terminal, unbranched or branched; flower pairs bracteate, pedunculate. Flowers bisexual, ebracteate, sessile. Tepals free, not coherent; posterior tepal erect or recurved; lateral; anterior tepals revolute. Staminal filaments adnate to tepals; anthers free, ovate, apiculate. Hypogynous glands connate, anterior, fleshy, truncate. Fruit globose, sometimes umbonate; outer mesocarp thin, not succulent. Seed 1, wingless, fleshy.