Medium to tall trees. Leaves opposite and decussate, imparipinnate, up to 6-jugate, with lateral leaflets subopposite; without domatia. Stipules 4 per node, lateral, ± foliaceous with main vein asymmetrically placed, leaving 2 contiguous interpetiolar scars on either side between adjoining leaf bases. Inflorescences terminal or axillary, paniculate, many-flowered, with terete branches and small caducous bracts, the branching varying from decussate in major branches through subopposite to ± spirally arranged in ultimate branchlets. Flowers bisexual, (4-)5(-6)-merous (except gynoecium), shortly pedicellate. Sepals valvate, subcoriaceous. Petals shorter than sepals, membranaceous, cuneate, bifid, with small glands at the apices. Stamens with subulate-terete filaments and oval to cordate anthers. Disc annular, upper surface ± smooth and with distinct impressions from filaments on outer edge or somewhat lobed. Ovary superior, syncarpous, 3(-4)-carpellate, each cell with a free subulate style and 2 or (3-)4 ovules. Fruit indehiscent, a 3(-4)-winged samara, obcordate in outline, the apex deeply incised, each wing formed by lateral expansion of the carpel wall; wings chartaceous with distinct reticulate venation; each locule usually with only a single seed, 1-3 seeds per fruit. Seeds oblong, the surface minutely papillate.