Gnetum gnemon L.

Buko (en)

Species

Gymnosperms > Gnetales > Gnetaceae > Gnetum

Characteristics

Tree or shrub, up to 22 m tall, 40 cm diam., without buttresses, exceptionally a climbing shrub. Crown monopodial, narrow, cylindrical; trunk grey, marked with conspicuous or faint rings. Leaves thin, yellow when dry, tapering at both ends, but varying in shape and size, 7½-20 by 2½-10 cm; secondary nerves bent, joining; petiole 6-18 mm. ♂ Inflorescences solitary, axillary, simple or once branched, yellowish, 3-6 cm long, collars 3 mm broad. ♂ Flowers with broad sporophyll, twice as long as the perianth (3 mm). Sterile ♀ flowers globose, tipped or beaked, 2 mm thick, 10-15 in a ring. ♀ Inflorescences similar. ♀ Flowers 5-8 at each node, globose, tipped or beaked, 3-4 mm long, inner tube exserted by 1 mm. Fruit ripening yellow, then orange-yellow or pink, sessile (exceptionally stalked), ellipsoid, shortly apiculate, 1-3½ cm long, almost velvety; middle envelop ribbed.
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Shrubs or small trees; bark grayish; crown narrow; branches becoming vinelike, green or yellowish green. Petiole 0.5-1.8 cm; leaf blade drying yellowish green, elliptic or oblong, 7.5-20 × 2.5-10 cm, leathery or membranous, lateral veins inconspicuous, curved, base attenuate into petiole, apex acuminate or cuspidate. Male inflorescences axillary, solitary, simple or once branched; male spikes (1-)3-6 cm × 2.5-3 mm, involucral collars clearly separated, to 1 cm apart, each collar with 50-80 flowers plus 5-15 globose sterile female flowers, basal hairs inconspicuous. Female inflorescences similar to male; nodes each with 5-8 female flowers. Seeds sessile or nearly so, yellow to orange-yellow or pink, ellipsoid, 1-2.5 (-3.5) × 0.9 (-1.5) cm, longitudinally ribbed, surface usually velvety, apex mucronate.
A shrub or small tree about 10 m high. It can grow to 20 m high. It normally has a cone shaped crown. The bark is greyish brown. The small branches are slender and like a vine. The leaves are produced in pairs opposite each other. They are dark green and shiny, oval and pointed at both ends. The leaves are 7.5-12.5 cm long by 2.5-7.5 cm wide. Trees are either male or female. Male spikes are 3-6 cm long. They consist of 50-80 very small flowers in rings along the stalk. Flowers are produced as cones made up of rings of scales along stalks 3-6 or 12 cm long. There are 5-8 flowers at each node. Fruits are oval and green when young but red when ripe. Fruit are 2-3 cm long and contain one seed.
Life form perennial
Growth form tree
Growth support free-standing
Foliage retention evergreen
Sexuality dioecy
Pollination entomogamy
Spread -
Mature width (meter) -
Mature height (meter) 19.0
Root system tap-root
Rooting depth (meter) -
Root diameter (meter) 0.3
Flower color -
Blooming months -
Fruit color
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

A tropical plant. It suits the lowland tropics. Trees occur in tropical rainforest from sea level to about 1200 metres altitude. It does best with rainfalls of 3000-5000 mm per year. It is best with an annual temperature of 22-30°C. It needs a temperature above 13°C. These trees grow in Malaysia, Indonesia and islands out to Fiji. They can grow on a range of soils but do best on deep well drained soils. It grows well in acid soils. It can grow in full sun but also tolerate shade. It cannot tolerate salt spray. It suits hardiness zones 10-12.
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Occasional in lowland ridges and mature fallow forest. Commonly found in secondary humid evergreen dipterocarp forests.
In rainforest at lower altitudes, but shrubby varieties ascending up to 1500 m in India.
Light 1-9
Soil humidity 4-6
Soil texture 1-4
Soil acidity -
Soil nutriment -
Hardiness (USDA) 9-12

Usage

Young leaf tips are eaten cooked. Young flowers are eaten, cooked. Young fruit are eaten, cooked. Ripe fruit are eaten raw or cooked. (Fruit should be crushed before cooking or they can explode.) Young flowers and fruit need cooking to get rid of irritating substances. The seeds are dried and flattened and then deep fried in hot oil and salt added.
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Uses. Young leaves and inflorescences are eaten as vegetable, fruits are also edible.
Uses animal food fiber fibre food fuel invertebrate food material medicinal oil poison social use wood
Edible flowers fruits leaves nuts saps seeds stems
Therapeutic use Antioxidants (leaf), Pelvic pain (leaf), Snake bites (leaf), Wounds and injuries (leaf)
Human toxicity -
Animal toxicity -

Cultivation

Trees are grown from seed. For using as seed for growing, a fully ripe red fruit is needed. Seeds exhibit a natural dormancy due to a hard impermeable seed coat. Breaking this seed coat can normally hasten germination. This is done by boring a hole into the seed. Propagation can be naturally by seeds distributed by birds, by planting seeds or by cuttings. A spacing of 6 m is suitable although in rows they are often put more closely. Trees appear to establish in the forest under-storey indicating some shade tolerance. Trees recover readily from pruning. Coppicing can produce a rapid flush of new leaves. Flushes of new leaves tends to occur seasonally. Trees can be topped to keep them shorter. If plants are grown by air layering they are shorter and more compact.
Mode cuttings seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Leaf

Gnetum gnemon leaf picture by Dieter Albrecht (cc-by-sa)

Fruit

Gnetum gnemon fruit picture by Dieter Albrecht (cc-by-sa)

Distribution

Gnetum gnemon world distribution map, present in China, Fiji, Hong Kong, Indonesia, India, Moldova (Republic of), Myanmar, Malaysia, Philippines, Solomon Islands, Thailand, and Viet Nam

Conservation status

Gnetum gnemon threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:383520-1
WFO ID wfo-0000795591
COL ID 6KN9L
BDTFX ID -
INPN ID -
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Gnetum gnemon Gnetum polystachyum Gnetum silvestris Gnetum silvestris Gnetum gnemon var. ovalifolium Gnetum gnemon var. sylvestris

Lower taxons

Gnetum gnemon var. brunonianum Gnetum gnemon var. griffithii Gnetum gnemon var. gnemon Gnetum gnemon var. tenerum Gnetum gnemon var. gracile